Huntington’s disease LIG1 modifier variant increases ligase fidelity and suppresses somatic CAG repeat expansion

E Eunhye Lee W Wonju Kim (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) D David H. Beier Y Yejin Lee (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) M Marina Kovalenko (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) F Faaiza Saif (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) E Esaria Oliver (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) B Bhairavi Srinageshwar (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) R Ryan Murtha (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) M Marissa A. Andrew (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) A Andrew Jiang (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) T Tammy Gillis (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) B Brigitte Demelo (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) J Jayla Ruliera (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) D Diane Lucente (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) S Seung Kwak (CHDI Management Inc.) R Ramee Lee (CHDI Management Inc.) R Ricardo Mouro Pinto (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) M Marcy E. MacDonald (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) J James F. Gusella (Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital) P Patrick J. O’Brien V Vanessa C. Wheeler I Ihn Sik Seong

Abstract

Huntington’s disease (HD) is a fatal neurodegenerative disorder caused by inheriting an expanded CAG repeat tract in the huntingtin gene ( HTT ) that further expands in somatic cells over an individual’s lifetime. Genome-wide association studies have provided critical insight into factors that modify the course of disease. These include DNA repair genes that alter the rate of somatic expansion and other genes that do not appear to directly influence this process. One modifier gene is DNA ligase 1 ( LIG1 ), in which a variant specifying a lysine to asparagine substitution (K845N) is associated with a profound (7 to 8 y) delay in the onset of motor signs. Here, we have taken a multifaceted approach to gain insight into the protective nature of this variant in HD. We demonstrate using in vitro ligase assays and enzyme kinetics that K845N enhances discrimination toward mismatched substrates and increases repair fidelity. Consistent with increased ligation fidelity, K845N confers protection against oxidative stress in cell-based assays. Finally, we demonstrate that the mouse LIG1 K843N orthologue suppresses somatic CAG expansion in HD knock-in mice. Overall, our data provide evidence that altered LIG1 function due to the K845N substitution may contribute to HD clinical delay by slowing somatic expansion in the brain and protecting the genome globally against damage. Significantly, our results provide a mechanistic foundation for considering DNA ligase fidelity as a therapeutic target in HD and potentially in other trinucleotide repeat disorders.

Article Details

Volume / Issue Vol. 123, Issue 10
Published March 10, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (23)

E

Eunhye Lee

W

Wonju Kim

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

D

David H. Beier

Y

Yejin Lee

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

M

Marina Kovalenko

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

F

Faaiza Saif

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

E

Esaria Oliver

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

B

Bhairavi Srinageshwar

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

R

Ryan Murtha

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

M

Marissa A. Andrew

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

A

Andrew Jiang

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

T

Tammy Gillis

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

B

Brigitte Demelo

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

J

Jayla Ruliera

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

D

Diane Lucente

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

S

Seung Kwak

CHDI Management Inc.

R

Ramee Lee

CHDI Management Inc.

R

Ricardo Mouro Pinto

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

M

Marcy E. MacDonald

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

J

James F. Gusella

Molecular Neurogenetics Unit, Center for Genomic Medicine, Massachusetts General Hospital

P

Patrick J. O’Brien

V

Vanessa C. Wheeler

I

Ihn Sik Seong