The importance of small-island populations for the long-term survival of endangered large-bodied insular mammals

S Sabhrina Gita Aninta (School of Biological and Behavioural Sciences, Queen Mary University of London) R Rosie Drinkwater A Alberto Carmagnini N Nicolas J. Deere (Durrell Institute of Conservation and Ecology, University of Kent) D Dwi Sendi Priyono (Department of Tropical Biology, Faculty of Biology, Universitas Gadjah Mada) N Noviar Andayani (Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Indonesia) N Nurul L. Winarni (Faculty of Mathematics and Natural Sciences, Research Center for Climate Change, Universitas Indonesia) J Jatna Supriatna M Matteo Fumagalli (School of Biological and Behavioural Sciences, Queen Mary University of London) G Greger Larson P Peter H. A. Galbusera (Antwerp Zoo Centre for Research and Conservation, Royal Zoological Society of Antwerp) A Alastair Macdonald (Royal (Dick) School of Veterinary Studies, University of Edinburgh) D Deborah Greer (School of Biological and Behavioural Sciences, Queen Mary University of London) K Kusdiantoro Mohamad (Division of Anatomy Histology and Embryology, School of Veterinary Medicine and Biomedical Sciences, Institut Pertanian Bogor (IPB) University) W Wahono Esthi Prasetyaningtyas (Division of Anatomy Histology and Embryology, School of Veterinary Medicine and Biomedical Sciences, Institut Pertanian Bogor (IPB) University) A Abdul Haris Mustari (Department of Forest Resources Conservation, Faculty of Forestry and Environment, Institut Pertanian Bogor (IPB) University) J John Lewis Williams (Department of Animal Science, Food and Nutrition, Università Cattolica del Sacro Cuore) R Ross Barnett (Durham Evolution and Ancient DNA, Department of Archaeology, University of Durham) D Darren Shaw (Royal (Dick) School of Veterinary Studies, University of Edinburgh) G Gono Semiadi (Research Centre for Biosystematics and Evolution, National Research and Innovation Agency (Badan Riset dan Inovasi Nasional)) J James Burton (International Union for Conservation of Nature and Natural Resources, Species Survival Commission (IUCN SSC) Asian Wild Cattle Specialist Group) D David J. I. Seaman (Durrell Institute of Conservation and Ecology, University of Kent) M Maria Voigt (Durrell Institute of Conservation and Ecology, University of Kent) M Matthew J. Struebig (Durrell Institute of Conservation and Ecology, University of Kent) S Selina Brace S Stephen J. Rossiter L Laurent Frantz

Abstract

Island populations of large vertebrates have experienced higher extinction rates than mainland populations over long timescales due to demographic stochasticity, genetic drift, and inbreeding. While being more susceptible to extinction and as such potentially targeted for conservation interventions such as genetic rescue, small-island populations can experience relatively less anthropogenic habitat degradation than those on larger islands. Here, we determine the consequences and conservation implications of long-term isolation and recent human activities on genetic diversity of island populations of two forest-dependent mammals endemic to the Wallacea archipelago: the anoa ( Bubalus spp.) and babirusa ( Babyrousa spp.). Using genomic analyses and habitat suitability models, we show that, compared to closely related species, populations on mainland Sulawesi exhibit low heterozygosity, high inbreeding, a high proportion of deleterious alleles, and experience a high rate of anthropogenic disturbance. In contrast, populations on smaller islands occupy higher-quality habitats, possess fewer deleterious mutations despite exhibiting lower heterozygosity and higher inbreeding. Site frequency spectra indicate that these patterns reflect stronger, long-term purging in smaller-island populations. Our results thus suggest that conservation efforts should focus on protecting small-island high-quality habitats and avoiding translocations from mainland populations. This study highlights the crucial role of small offshore islands for the long-term survival of Wallacea’s iconic and indigenous mammals in the face of development on the mainland.

Article Details

Volume / Issue Vol. 122, Issue 26
Published July 01, 2025
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (27)

S

Sabhrina Gita Aninta

School of Biological and Behavioural Sciences, Queen Mary University of London

R

Rosie Drinkwater

A

Alberto Carmagnini

N

Nicolas J. Deere

Durrell Institute of Conservation and Ecology, University of Kent

D

Dwi Sendi Priyono

Department of Tropical Biology, Faculty of Biology, Universitas Gadjah Mada

N

Noviar Andayani

Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Indonesia

N

Nurul L. Winarni

Faculty of Mathematics and Natural Sciences, Research Center for Climate Change, Universitas Indonesia

J

Jatna Supriatna

M

Matteo Fumagalli

School of Biological and Behavioural Sciences, Queen Mary University of London

G

Greger Larson

P

Peter H. A. Galbusera

Antwerp Zoo Centre for Research and Conservation, Royal Zoological Society of Antwerp

A

Alastair Macdonald

Royal (Dick) School of Veterinary Studies, University of Edinburgh

D

Deborah Greer

School of Biological and Behavioural Sciences, Queen Mary University of London

K

Kusdiantoro Mohamad

Division of Anatomy Histology and Embryology, School of Veterinary Medicine and Biomedical Sciences, Institut Pertanian Bogor (IPB) University

W

Wahono Esthi Prasetyaningtyas

Division of Anatomy Histology and Embryology, School of Veterinary Medicine and Biomedical Sciences, Institut Pertanian Bogor (IPB) University

A

Abdul Haris Mustari

Department of Forest Resources Conservation, Faculty of Forestry and Environment, Institut Pertanian Bogor (IPB) University

J

John Lewis Williams

Department of Animal Science, Food and Nutrition, Università Cattolica del Sacro Cuore

R

Ross Barnett

Durham Evolution and Ancient DNA, Department of Archaeology, University of Durham

D

Darren Shaw

Royal (Dick) School of Veterinary Studies, University of Edinburgh

G

Gono Semiadi

Research Centre for Biosystematics and Evolution, National Research and Innovation Agency (Badan Riset dan Inovasi Nasional)

J

James Burton

International Union for Conservation of Nature and Natural Resources, Species Survival Commission (IUCN SSC) Asian Wild Cattle Specialist Group

D

David J. I. Seaman

Durrell Institute of Conservation and Ecology, University of Kent

M

Maria Voigt

Durrell Institute of Conservation and Ecology, University of Kent

M

Matthew J. Struebig

Durrell Institute of Conservation and Ecology, University of Kent

S

Selina Brace

S

Stephen J. Rossiter

L

Laurent Frantz