The global diversity and decline of glacier animals

A Andrea Simoncini (Department of Environmental Science and Policy, University of Milan) I Isabel Cantera (Department of Environmental Science and Policy, University of Milan) S Simone Giachello (Department of Environmental Science and Policy, University of Milan) B Barbara Valle (Department of Life Sciences, University of Siena) M Mauro Gobbi (Research and Museum Collections Office, Climate and Ecology Unit, MUSE - Science Museum) D Diego Fontaneto (National Biodiversity Future Center) K Krzysztof Zawierucha (Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University) R Ronald Laniecki (Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University) S Stephen James Coulson (Department of Arctic Biology, The University Centre in Svalbard, Longyearbyen) V Valeria Lencioni (Research and Museum Collections Office, Climate and Ecology Unit, MUSE - Science Museum) S Sophie Cauvy-Fraunié (French National Research Institute for Agriculture, Food, and Environment, Unité de Recherche RiverLy, Centre de Lyon-Villeurbanne) A Alejandra Morán Ordóñez (Department of Environmental Sciences, Landscape Ecology and Ecosystems Conservation unit, University of Basel) G Gianalberto Losapio (Faculty of Geosciences and Environment, Institute of Earth Surface Dynamics, University of Lausanne) N Nozomu Takeuchi (Department of Earth Sciences, Center for Environmental Remote Sensing, Chiba University) Q Qiao Liu D Daniel H. Shain (Biology Department, Rutgers The State University of New Jersey) H Heather Fair (Institute of Mountain Hazards and Environment, Chinese Academy of Sciences) K Karel Janko (Laboratory of Non-Mendelian Evolution, Institute of Animal Physiology and Genetics AS CR) M Miloslav Devetter (Institute of Soil Biology and Biogeochemistry, Biology Centre of the Czech Academy of Sciences) M Marco Antonio Jiménez-Santos (Laboratory of Non-Mendelian Evolution, Institute of Animal Physiology and Genetics AS CR) M Marie Šabacká (Department of Ecology, Faculty of Natural Sciences, Charles University in Prague) M Mattia Falaschi (Department of Environmental Science and Policy, University of Milan) S Silvio Marta (Institute of Geosciences and Earth Resources, Italian National Research Council) G Gentile Francesco Ficetola (Department of Environmental Science and Policy, University of Milan)

Abstract

Glacier environments host an unrecognized number of animals that are at high risk of extinction due to global warming. Yet, the absence of comprehensive analyses has hampered the development of a coordinated agenda for research and conservation. By assembling a global dataset of terrestrial and freshwater animals in glacial habitats, we show that glacier environments host at least 152 animal species from 14 classes and 7 phyla. This diversity is underestimated for nearly all the groups and we detected widespread taxonomical, ecological, and geographical biases. Glacier animals exhibited diverse ecological patterns, with 73 species reported exclusively from glacial habitats (glacier specialists). Biogeographical patterns showed clear differences between groups, with some taxa (tardigrades, rotifers) found mostly in polar areas and others (insects, springtails) associated with tropical and temperate mountains. Glacier regions with a lower temperature showed a higher richness of glacier specialists. Glacier animals were widespread across the metazoan tree of life; the ability to disperse passively by wind emerged as the strongest driver of the occurrence of glacier animals. By linking the distribution of glacier specialists to scenarios of glacier retreat, we identified areas and species that are likely to experience abrupt declines in the next few decades, as well as species that could face complete habitat loss. Our results call for urgent actions to secure the future of glacier environments and their singular, understudied fauna.

Article Details

Volume / Issue Vol. 123, Issue 25
Published June 23, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (24)

A

Andrea Simoncini

Department of Environmental Science and Policy, University of Milan

I

Isabel Cantera

Department of Environmental Science and Policy, University of Milan

S

Simone Giachello

Department of Environmental Science and Policy, University of Milan

B

Barbara Valle

Department of Life Sciences, University of Siena

M

Mauro Gobbi

Research and Museum Collections Office, Climate and Ecology Unit, MUSE - Science Museum

D

Diego Fontaneto

National Biodiversity Future Center

K

Krzysztof Zawierucha

Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University

R

Ronald Laniecki

Department of Animal Taxonomy and Ecology, Faculty of Biology, Adam Mickiewicz University

S

Stephen James Coulson

Department of Arctic Biology, The University Centre in Svalbard, Longyearbyen

V

Valeria Lencioni

Research and Museum Collections Office, Climate and Ecology Unit, MUSE - Science Museum

S

Sophie Cauvy-Fraunié

French National Research Institute for Agriculture, Food, and Environment, Unité de Recherche RiverLy, Centre de Lyon-Villeurbanne

A

Alejandra Morán Ordóñez

Department of Environmental Sciences, Landscape Ecology and Ecosystems Conservation unit, University of Basel

G

Gianalberto Losapio

Faculty of Geosciences and Environment, Institute of Earth Surface Dynamics, University of Lausanne

N

Nozomu Takeuchi

Department of Earth Sciences, Center for Environmental Remote Sensing, Chiba University

Q

Qiao Liu

D

Daniel H. Shain

Biology Department, Rutgers The State University of New Jersey

H

Heather Fair

Institute of Mountain Hazards and Environment, Chinese Academy of Sciences

K

Karel Janko

Laboratory of Non-Mendelian Evolution, Institute of Animal Physiology and Genetics AS CR

M

Miloslav Devetter

Institute of Soil Biology and Biogeochemistry, Biology Centre of the Czech Academy of Sciences

M

Marco Antonio Jiménez-Santos

Laboratory of Non-Mendelian Evolution, Institute of Animal Physiology and Genetics AS CR

M

Marie Šabacká

Department of Ecology, Faculty of Natural Sciences, Charles University in Prague

M

Mattia Falaschi

Department of Environmental Science and Policy, University of Milan

S

Silvio Marta

Institute of Geosciences and Earth Resources, Italian National Research Council

G

Gentile Francesco Ficetola

Department of Environmental Science and Policy, University of Milan