Temperature and developmental stage govern intestinal susceptibility to human coronavirus 229E

A Aleksandra Synowiec (Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University) L Laurensius Kevin Lie (Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University) K Katarzyna Owczarek N Nina Johannesson (OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam) N Nina Mickiewicz (Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University) H Heng-Chang Chen (Quantitative Virology Research Group, Population Diagnostics Center, Łukasiewicz Research Network–PORT Polish Center for Technology Development) A Artur Szczepański (Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University) M Madison S. Strine R Renata B. Filler (Department of Laboratory Medicine, Yale School of Medicine) M Maciej Borowiec (Department of Clinical and Laboratory Genetics, Medical University of Lodz) M Michał Pietrusiński (Department of Clinical and Laboratory Genetics, Medical University of Lodz) I Izabela Dróżdż (Department of Clinical and Laboratory Genetics, Medical University of Lodz) A Agnieszka Robaszkiewicz (Department of General Biophysics, Faculty of Biology and Environmental Protection, University of Lodz) M Michał Bochenek (Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University) M Matthias Zilbauer (Department of Pediatrics, School of Clinical Medicine, University of Cambridge) J Justyna Rymarowicz (Department of General Surgery, Jagiellonian University Medical College) M Michał Pędziwiatr (Department of General Surgery, Jagiellonian University Medical College) L Liza Konnikova (Pediatrics Department, Yale University School of Medicine) C Craig B. Wilen (Department of Laboratory Medicine, Yale School of Medicine) D Dasja Pajkrt (OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam) K Katja C. Wolthers (OrganoVIR Labs, Department of Medical Microbiology, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, University of Amsterdam) C Carlemi Calitz (OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam) A Adithya Sridhar (OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam) K Krzysztof Pyrc

Abstract

Human coronaviruses have been primarily associated with upper respiratory tract infections, yet cases of gastrointestinal symptoms in COVID-19 patients have highlighted their potential to cause systemic disease. Here, we detail the infection of intestinal epithelia by an endemic, low-pathogenic human coronavirus, human alphacoronavirus 229E, using patient-derived human intestinal enteroids (HIEs) from donors of various ages. Using fetal, pediatric, and adult HIEs, we investigated how physiologically relevant temperatures: 37 °C and 32 °C, reflecting gastrointestinal and upper-airway conditions, respectively, modulate epithelial responses and viral infection dynamics. We show that there is temperature-dependent transcriptional reprogramming, indicating strong temperature-dependent regulation of virus replication and epithelial responses. Among the seasonal coronaviruses tested, only HCoV-229E productively infects HIEs. At 32 °C, HCoV-229E replicates efficiently in enteroids from all donor ages and releases high titers of infectious progeny. In contrast, at 37 °C, productive replication is largely confined to fetal and a subset of pediatric tissues, revealing a developmental and temperature-sensitive restriction on infection. Confocal and flow cytometry analyses identify enterocytes as the primary target cells for HCoV-229E. Furthermore, we show that camostat, a serine protease inhibitor, significantly reduces HCoV-229E replication in HIEs, confirming a critical role for host serine protease activity. Collectively, these findings establish HIEs as a relevant model for HCoV-229E–host interactions and reveal temperature- and age-dependent determinants governing intestinal permissiveness to this seasonal coronavirus.

Article Details

Volume / Issue Vol. 123, Issue 26
Published June 30, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (24)

A

Aleksandra Synowiec

Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University

L

Laurensius Kevin Lie

Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University

K

Katarzyna Owczarek

N

Nina Johannesson

OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam

N

Nina Mickiewicz

Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University

H

Heng-Chang Chen

Quantitative Virology Research Group, Population Diagnostics Center, Łukasiewicz Research Network–PORT Polish Center for Technology Development

A

Artur Szczepański

Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University

M

Madison S. Strine

R

Renata B. Filler

Department of Laboratory Medicine, Yale School of Medicine

M

Maciej Borowiec

Department of Clinical and Laboratory Genetics, Medical University of Lodz

M

Michał Pietrusiński

Department of Clinical and Laboratory Genetics, Medical University of Lodz

I

Izabela Dróżdż

Department of Clinical and Laboratory Genetics, Medical University of Lodz

A

Agnieszka Robaszkiewicz

Department of General Biophysics, Faculty of Biology and Environmental Protection, University of Lodz

M

Michał Bochenek

Virogenetics Laboratory of Virology, Małopolska Centre of Biotechnology, Jagiellonian University

M

Matthias Zilbauer

Department of Pediatrics, School of Clinical Medicine, University of Cambridge

J

Justyna Rymarowicz

Department of General Surgery, Jagiellonian University Medical College

M

Michał Pędziwiatr

Department of General Surgery, Jagiellonian University Medical College

L

Liza Konnikova

Pediatrics Department, Yale University School of Medicine

C

Craig B. Wilen

Department of Laboratory Medicine, Yale School of Medicine

D

Dasja Pajkrt

OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam

K

Katja C. Wolthers

OrganoVIR Labs, Department of Medical Microbiology, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, University of Amsterdam

C

Carlemi Calitz

OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam

A

Adithya Sridhar

OrganoVIR Labs, Emma Children’s Hospital, Department of Pediatric Infectious Diseases, Amsterdam University Medical Center, Academic Medical Center, Amsterdam Institute for Infection and Immunity, Amsterdam Institute for Reproduction and Development, University of Amsterdam

K

Krzysztof Pyrc