TCA cycle mode switch determines the fate of pirtobrutinib-tolerant persister cells in mantle cell lymphoma

W Wei Wang Q Qingsong Cai (1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX) Y Yang Liu L Lei Nie (Department of Oncology Shanxi Provincial Cancer Hospital Xian China) H Heng-Huan Lee F Fangfang Yan Y Yue Fei Y Yixin Yao Y Yijing Li (College of Mining, Liaoning Technical University 1 , Fuxin 123000, Liaoning,) L Lin Tan P Philip L. Lorenzi Y Ying-Nai Wang J Jun Yao (Key Lab of Mesoscopic Chemistry, School of Chemistry and Chemical Engineering) Z Zhihong Chen J Joseph Mitchell McIntosh (1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX) C Cheng-Tai Yu (1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX) P Preetesh Jain V Vivian C. Jiang (1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX) J Jovanny Vargas (1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX) X Xiaolin Li (Energy and Environmental Directorate, Pacific Northwest National Laboratory) T Tianci Zhang S Shaoying Li D David Santos S Selvi Thirumurthi E Erin Heather Seeley (7Department of Chemistry, Mass Spectrometry Imaging Facility at The University of Texas at Austin, Austin, TX) L Lukas Mikolaj Simon (8Verna and Marrs McLean Department of Biochemistry and Molecular Pharmacology, Therapeutic Innovation Center at Baylor College of Medicine, Houston, TX) C Christopher Flowers (1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX) C Chi Young Ok (4Division of Pathology and Laboratory Medicine, Department of Hematopathology, The University of Texas MD Anderson Cancer Center, Houston, TX) M Michael Wang

Abstract

Abstract Bruton tyrosine kinase inhibitors (BTKis) and cell therapy have successfully been used to treat mantle cell lymphoma (MCL). However, therapy resistance inevitably emerges. Cancer cells can progressively develop stable resistance by traversing through a transient drug-tolerant persister (DTP) state. The mechanisms enabling DTP cells to reversibly adapt to therapies and evolve to acquire heterogeneity remain poorly understood, and characterizing DTP cells in MCL continues to pose a challenge for clinic translation. Here, using pirtobrutinib, a recently US Food and Drug Administration–approved noncovalent BTKi, we identified pirtobrutinib-tolerant persister cells exhibiting morphological variability by presenting a unique population of enlarged cells (giant cells) with reversible fate transitions. During treatment, giant cells enter a nonproliferative, dedifferentiated state, addicted to an activated cytosolic tricarboxylic acid (TCA) cycle coupled with the malate-aspartate shuttle to engage in biosynthesis. Upon drug removal, the TCA cycle shifts to oxidative catabolism, promoting giant cells to differentiate into regular-sized cells. Throughout the transition, acetyl coenzyme A modulates cell fate by fine-tuning stemness. Our biphasic model demonstrates that the metabolic switch governs the phenotypic plasticity of DTP cells in MCL, resulting in a dynamic presence of DTP cells across various developmental states in response to systemic therapies. Targeting giant cells before their differentiation offers a promising strategy to overcoming therapy resistance in MCL.

Article Details

Journal Blood
Volume / Issue Vol. 146, Issue 21
Published November 20, 2025
Pages 2544-2560
ISSN 0006-4971
Publisher Elsevier BV

Journal Info

Blood

Elsevier BV

ISSN: 0006-4971 Health Sciences

Authors (29)

W

Wei Wang

Q

Qingsong Cai

1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX

Y

Yang Liu

L

Lei Nie

Department of Oncology Shanxi Provincial Cancer Hospital Xian China

H

Heng-Huan Lee

F

Fangfang Yan

Y

Yue Fei

Y

Yixin Yao

Y

Yijing Li

College of Mining, Liaoning Technical University 1 , Fuxin 123000, Liaoning,

L

Lin Tan

P

Philip L. Lorenzi

Y

Ying-Nai Wang

J

Jun Yao

Key Lab of Mesoscopic Chemistry, School of Chemistry and Chemical Engineering

Z

Zhihong Chen

J

Joseph Mitchell McIntosh

1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX

C

Cheng-Tai Yu

1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX

P

Preetesh Jain

V

Vivian C. Jiang

1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX

J

Jovanny Vargas

1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX

X

Xiaolin Li

Energy and Environmental Directorate, Pacific Northwest National Laboratory

T

Tianci Zhang

S

Shaoying Li

D

David Santos

S

Selvi Thirumurthi

E

Erin Heather Seeley

7Department of Chemistry, Mass Spectrometry Imaging Facility at The University of Texas at Austin, Austin, TX

L

Lukas Mikolaj Simon

8Verna and Marrs McLean Department of Biochemistry and Molecular Pharmacology, Therapeutic Innovation Center at Baylor College of Medicine, Houston, TX

C

Christopher Flowers

1Department of Lymphoma and Myeloma, The University of Texas MD Anderson Cancer Center, Houston, TX

C

Chi Young Ok

4Division of Pathology and Laboratory Medicine, Department of Hematopathology, The University of Texas MD Anderson Cancer Center, Houston, TX

M

Michael Wang