SARS-CoV-2 S assembly into virions facilitated by host ERM proteins

J Jiaming Wang (School of Life Sciences, Beijing University of Chinese Medicine) W Wanbo Tai (New Cornerstone Science Laboratory, Tsinghua University-Peking University, Joint Center for Life Sciences, School of Basic Medical Sciences, Tsinghua University) Z Zhaoyang Wang W Wenxin Dai (School of Life Sciences, Beijing University of Chinese Medicine) M Mingrui Yang (School of Life Sciences, Beijing University of Chinese Medicine) J Jiajian Guo (School of Life Sciences, Beijing University of Chinese Medicine) P Pengfei He (School of Life Sciences, Beijing University of Chinese Medicine) Y Yanan Nan (School of Life Sciences, Beijing University of Chinese Medicine) T Tianyu Li S Shuqi Zhou (Tsinghua University , , ,) D Dongxiao Cui (School of Life Sciences, Beijing University of Chinese Medicine) Y Yiqun Li (School of Life Sciences, Beijing University of Chinese Medicine) C Cuiyan Ma (School of Life Sciences, Beijing University of Chinese Medicine) Y Yue Zhang D Dongdong Li (Key Laboratory for Ultrafine Materials of Ministry of Education, Shanghai Engineering Research Center of Hierarchical Nanomaterials, School of Materials Science and Engineering, East China University of Science and Technology, 130 Meilong Road, Shanghai 200237, China) Z Zhengdan Zhu (DP Technology) K Kexin Chu (DP Technology) D Dongdong Wang S Songhui Yang (Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, State Key Laboratory of Pathogen and Biosecurity, Key Laboratory of Jilin Province for Zoonosis Prevention and Control) X Xinyu Zhuang (Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, State Key Laboratory of Pathogen and Biosecurity, Key Laboratory of Jilin Province for Zoonosis Prevention and Control) M Mingyao Tian (Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, State Key Laboratory of Pathogen and Biosecurity, Key Laboratory of Jilin Province for Zoonosis Prevention and Control) M Mingkang Huang (Institute of Infectious Diseases, Shenzhen Bay Laboratory) X Xianwen Zhang G Gong Cheng W Wenfu Ma (School of Life Sciences, Beijing University of Chinese Medicine)

Abstract

The host cell cytoskeleton plays a critical role in the SARS-CoV-2 life cycle, though the underlying mechanisms remain poorly understood. This study investigates the interaction between the SARS-CoV-2 spike (S) protein and the cytoskeleton-associated ezrin-radixin-moesin (ERM) proteins through biochemical and structural characterization. A previously unidentified ERM-binding motif on the SARS-CoV-2 S protein is identified, revealing that S-ERM interactions are specifically conserved among highly pathogenic coronaviruses, including SARS-CoV, MERS-CoV, and SARS-CoV-2. Functionally, these interactions facilitate S packaging into virions by directing it to assembly sites, utilizing ERM’s affinity for negatively curved membranes, akin to its role in cell surface protrusions. Silencing ERM expression significantly reduces SARS-CoV-2 titer, highlighting its essential role in viral propagation. Additionally, leveraging the established role of COPI-mediated trafficking in S localization, a compound is developed to disrupt S-COPI binding, promoting S secretion to the cell surface and effectively reducing viral titers. Our findings revealed a critical host–pathogen interaction that drives S incorporation into virions and identified ERM proteins as key facilitators of coronavirus assembly. Furthermore, our study suggests an antiviral strategy by targeting the S-COPI trafficking pathway. These insights advanced our understanding of coronavirus–host interactions and provided a potential therapeutic approach against SARS-CoV-2 and other highly pathogenic coronaviruses.

Article Details

Volume / Issue Vol. 123, Issue 5
Published February 03, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (25)

J

Jiaming Wang

School of Life Sciences, Beijing University of Chinese Medicine

W

Wanbo Tai

New Cornerstone Science Laboratory, Tsinghua University-Peking University, Joint Center for Life Sciences, School of Basic Medical Sciences, Tsinghua University

Z

Zhaoyang Wang

W

Wenxin Dai

School of Life Sciences, Beijing University of Chinese Medicine

M

Mingrui Yang

School of Life Sciences, Beijing University of Chinese Medicine

J

Jiajian Guo

School of Life Sciences, Beijing University of Chinese Medicine

P

Pengfei He

School of Life Sciences, Beijing University of Chinese Medicine

Y

Yanan Nan

School of Life Sciences, Beijing University of Chinese Medicine

T

Tianyu Li

S

Shuqi Zhou

Tsinghua University , , ,

D

Dongxiao Cui

School of Life Sciences, Beijing University of Chinese Medicine

Y

Yiqun Li

School of Life Sciences, Beijing University of Chinese Medicine

C

Cuiyan Ma

School of Life Sciences, Beijing University of Chinese Medicine

Y

Yue Zhang

D

Dongdong Li

Key Laboratory for Ultrafine Materials of Ministry of Education, Shanghai Engineering Research Center of Hierarchical Nanomaterials, School of Materials Science and Engineering, East China University of Science and Technology, 130 Meilong Road, Shanghai 200237, China

Z

Zhengdan Zhu

DP Technology

K

Kexin Chu

DP Technology

D

Dongdong Wang

S

Songhui Yang

Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, State Key Laboratory of Pathogen and Biosecurity, Key Laboratory of Jilin Province for Zoonosis Prevention and Control

X

Xinyu Zhuang

Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, State Key Laboratory of Pathogen and Biosecurity, Key Laboratory of Jilin Province for Zoonosis Prevention and Control

M

Mingyao Tian

Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, State Key Laboratory of Pathogen and Biosecurity, Key Laboratory of Jilin Province for Zoonosis Prevention and Control

M

Mingkang Huang

Institute of Infectious Diseases, Shenzhen Bay Laboratory

X

Xianwen Zhang

G

Gong Cheng

W

Wenfu Ma

School of Life Sciences, Beijing University of Chinese Medicine