Sanger Sequencing of Xeno‐Nucleic Acid

Y Yueyao Wang Z Ze Zhang (Department of Polymer Science and Engineering) L Lekang Chen (State Key Laboratory of Coordination Chemistry Department of Biomedical Engineering College of Engineering and Applied Sciences Chemistry and Biomedicine Innovation Center (ChemBIC) Jiangsu Key Laboratory of Artificial Functional Materials Nanjing University Nanjing Jiangsu 210023 China) J Jing Zhu (Hefei National Research Center for Physical Sciences at the Microscale, CAS Key Laboratory of Strongly-Coupled Quantum Matter Physics, Key Laboratory of Surface and Interface Chemistry and Energy Catalysis of Anhui Higher Education Institutes, Department of Chemical Physics) S Shunfu Qin (School of Pharmacy Nanjing University of Chinese Medicine Nanjing Jiangsu 210023 China) D Dongying Wei (State Key Laboratory of Coordination Chemistry Department of Biomedical Engineering College of Engineering and Applied Sciences Chemistry and Biomedicine Innovation Center (ChemBIC) Jiangsu Key Laboratory of Artificial Functional Materials Nanjing University Nanjing Jiangsu 210023 China) J Jibao Zhang (Nanjing Superyears Gene Technology Co., Ltd. Nanjing Jiangsu 211800 China) F Feng Han (State Key Laboratory of Coordination Chemistry, School of Chemistry and Chemical Engineering, Chemistry and Biomedicine Innovation Center (ChemBIC), ChemBioMed Interdisciplinary Research Center at Nanjing University) F Fulong Wang Q Qi Lin (Department of Chemistry) R Rui Fan X Xintong Li (Shanghai Key Laboratory of Green Chemistry and Chemical Processes, State Key Laboratory of Petroleum Molecular & Process Engineering, School of Chemistry and Molecular Engineering, East China Normal University, North Zhongshan Rd. 3663, Shanghai 200062, China) H Hanyang Yu (State Key Laboratory of Coordination Chemistry, Department of Biomedical Engineering, College of Engineering and Applied Sciences, Chemistry and Biomedicine Innovation Center (ChemBIC), ChemBioMed Interdisciplinary Research Center at Nanjing University)

Abstract

Abstract Xeno‐nucleic acids (XNAs) offer biostable genetic polymers for biotechnology and information technology, but are incompatible with prevailing DNA sequencing methodologies. Here we report a Sanger sequencing approach tailored for XNA, enabling direct readout of XNA strands up to 50 bases. This new method relies on a Bst DNA polymerase mutant, which not only recognizes XNA as a template but also efficiently incorporates dideoxyribonucleoside triphosphate (ddNTP) substrates, facilitating sequencing of XNA by synthesis of complementary DNA. The Bst F710Y mutant exhibits at least two orders of magnitude higher activity in catalyzing ddNTP incorporation. Our method demonstrates accurate sequence determination of diverse XNA strands of distinct backbone chemistries such as TNA and FANA. Lastly, we showcase proof‐of‐concept automated XNA sequencing on a genetic analyzer instrument, by leveraging Bst mutant's ability to stochastically incorporate BigDye‐labeled ddNTP substrates in a single reaction. This work establishes a direct sequencing platform for XNA employing a chain termination strategy, and lays a foundation for future de novo identification of functional XNA and development of XNA‐based data storage system.

Article Details

Volume / Issue Vol. 65, Issue 9
Published February 23, 2026
ISSN 1433-7851
Publisher Wiley

Journal Info

Angewandte Chemie International Edition

Wiley

ISSN: 1433-7851 Physical Sciences

Authors (13)

Y

Yueyao Wang

Z

Ze Zhang

Department of Polymer Science and Engineering

L

Lekang Chen

State Key Laboratory of Coordination Chemistry Department of Biomedical Engineering College of Engineering and Applied Sciences Chemistry and Biomedicine Innovation Center (ChemBIC) Jiangsu Key Laboratory of Artificial Functional Materials Nanjing University Nanjing Jiangsu 210023 China

J

Jing Zhu

Hefei National Research Center for Physical Sciences at the Microscale, CAS Key Laboratory of Strongly-Coupled Quantum Matter Physics, Key Laboratory of Surface and Interface Chemistry and Energy Catalysis of Anhui Higher Education Institutes, Department of Chemical Physics

S

Shunfu Qin

School of Pharmacy Nanjing University of Chinese Medicine Nanjing Jiangsu 210023 China

D

Dongying Wei

State Key Laboratory of Coordination Chemistry Department of Biomedical Engineering College of Engineering and Applied Sciences Chemistry and Biomedicine Innovation Center (ChemBIC) Jiangsu Key Laboratory of Artificial Functional Materials Nanjing University Nanjing Jiangsu 210023 China

J

Jibao Zhang

Nanjing Superyears Gene Technology Co., Ltd. Nanjing Jiangsu 211800 China

F

Feng Han

State Key Laboratory of Coordination Chemistry, School of Chemistry and Chemical Engineering, Chemistry and Biomedicine Innovation Center (ChemBIC), ChemBioMed Interdisciplinary Research Center at Nanjing University

F

Fulong Wang

Q

Qi Lin

Department of Chemistry

R

Rui Fan

X

Xintong Li

Shanghai Key Laboratory of Green Chemistry and Chemical Processes, State Key Laboratory of Petroleum Molecular & Process Engineering, School of Chemistry and Molecular Engineering, East China Normal University, North Zhongshan Rd. 3663, Shanghai 200062, China

H

Hanyang Yu

State Key Laboratory of Coordination Chemistry, Department of Biomedical Engineering, College of Engineering and Applied Sciences, Chemistry and Biomedicine Innovation Center (ChemBIC), ChemBioMed Interdisciplinary Research Center at Nanjing University