Reward prediction is encoded by orexin neuron activity during motivated behavior

Y Yutao Dong (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine) S Sheikh Mizanur Rahaman (Stress Recognition and Response, Research Institute of Environmental Medicine, Nagoya University) W Wenjun Zhu A Ayumu Inutsuka (Department of Physiology, Aichi Medical University) D Daisuke Ono (Stress Recognition and Response, Research Institute of Environmental Medicine, Nagoya University) R Rinako Tanaka (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine) T Tetsuo Matsuzaki (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine) E Eiji Shibata (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine) M Madoka Isobe (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine) S Shuntaro Izawa (Department of Neuronal Control of Metabolism, Max Planck Institute for Metabolism Research) A Akihiro Yamanaka (Chinese Institute for Brain Research) K Kiyofumi Yamada (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine) H Hiroyuki Mizoguchi (Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine)

Abstract

Orexin neurons regulate physiological functions, such as energy homeostasis, wakefulness, and motivated behaviors. However, studies linking orexin neuron activity to behavior via selectively activating/inactivating these inputs in a temporally controlled manner in rats are scarce. Here, we examined the role that orexin neurons play in motivated behavior in transgenic rats using cell type–specific fiber photometry and optogenetic manipulation. Using chemogenetics, we found that motivation for a reward increased when orexin neurons were activated. Furthermore, during motivated behavior, orexin neuron activity changed dynamically: Activity increased during reward prediction and decreased after reward receipt. When an unexpected event occurred (i.e., not obtaining an expected reward), increased orexin activity was sustained. Notably, orexin activity strengthened with increasing effort. Optogenetic inhibition of orexin neuron activation during reward prediction and treatment with an orexin 1-receptor antagonist reduced reward-seeking behavior. Therefore, orexin is crucial for linking predicted expectations with motivated behavior. Moreover, optimization of orexin activity is necessary to overcome difficulties during motivated behaviors.

Article Details

Volume / Issue Vol. 123, Issue 27
Published July 07, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (13)

Y

Yutao Dong

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine

S

Sheikh Mizanur Rahaman

Stress Recognition and Response, Research Institute of Environmental Medicine, Nagoya University

W

Wenjun Zhu

A

Ayumu Inutsuka

Department of Physiology, Aichi Medical University

D

Daisuke Ono

Stress Recognition and Response, Research Institute of Environmental Medicine, Nagoya University

R

Rinako Tanaka

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine

T

Tetsuo Matsuzaki

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine

E

Eiji Shibata

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine

M

Madoka Isobe

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine

S

Shuntaro Izawa

Department of Neuronal Control of Metabolism, Max Planck Institute for Metabolism Research

A

Akihiro Yamanaka

Chinese Institute for Brain Research

K

Kiyofumi Yamada

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine

H

Hiroyuki Mizoguchi

Department of Neuropsychopharmacology and Hospital Pharmacy, Nagoya University Graduate School of Medicine