Reprogramming glutamine metabolism enhances <i>BCMA</i> -CAR T-cell fitness and therapeutic efficacy in multiple myeloma

F Flor Navarro (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) T Teresa Lozano (Centro de Investigación Médica Aplicada, CIMA, Pamplona, Spain) A Andrea Fuentes-García (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) I Inés Sánchez-Moreno (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) M Marta Larrayoz P Pedro Justicia (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) B Beatriz Perucha (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) M Maialen Martinez-Tabar (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) R Rebeca Martinez-Turrillas (2Hemato-Oncology Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) N Noelia Casares (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) C Celia Martín-Otal M Marta Gorraiz (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) E Erin W. Meermeier (1Department of Immunology, Mayo Clinic Arizona, Scottsdale, AZ) M Marta Chesi D Douglas Lake (5School of Life Sciences, Arizona State University, Phoenix, AZ) P P. Leif Bergsagel (Mayo Clinic Arizona, Scottsdale) E Eva Santamaría (6RNA Biology and Therapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, University of Navarra, Pamplona, Spain) M Maria Erendira Calleja-Cervantes P Patxi San Martín-Uriz (1Center for Applied Medical Research, Universidad de Navarra, Navarra Medical Research Institute, Pamplona, Spain) L Lorea Jordana-Urriza (2Hemato-Oncology Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) X Xabier Agirre S Sandra Hervás-Stubbs (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) J Juan Roberto Rodriguez-Madoz (1Center for Applied Medical Research, Universidad de Navarra, Navarra Medical Research Institute, Pamplona, Spain) J José Ángel Martinez-Climent (2Hemato-Oncology Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain) F Felipe Prosper J Juan José Lasarte (1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain)

Abstract

Abstract Glutamine dependence of cancer cells reduces local glutamine availability, which hinders antitumor T-cell functionality and facilitates immune evasion. We thus speculated that glutamine deprivation might be limiting efficacy of chimeric antigen receptor (CAR) T-cell therapies in patients with cancer. We have seen that antigen-specific T cells are unable to proliferate or produce interferon gamma (IFN-γ) in response to antigen stimulation when glutamine concentration is limited. Using multiple myeloma (MM) as a glutamine-dependent disease model, we found that murine CAR T cells selectively targeting B-cell maturation antigen (Bcma) in MM cells were sensitive to glutamine deprivation. However, CAR T cells engineered to increase glutamine uptake by expression of the glutamine transporter Asct2 exhibited enhanced proliferation and responsiveness to antigen stimulation, increased production of IFN-γ, and heightened cytotoxic activity, even under conditions of low glutamine concentration. Mechanistically, Asct2 overexpression reprogrammed the metabolic fitness of CAR T cells by upregulating the mechanistic target of rapamycin complex 1 gene signature, modifying the solute carrier transporter repertoire, and improving both basal oxygen consumption rate and glycolytic function, thereby enhancing CAR T-cell persistence in vivo. Accordingly, expression of Asct2 increased the efficacy of Bcma-CAR T cells in syngeneic and genetically engineered mouse models of MM, which prolonged mouse survival. In patients, higher-level expression of ASCT2 by MM cells predicted poor outcome to combined immunotherapy and BCMA-CAR T-cell therapy. Our results indicate that reprogramming glutamine metabolism may enhance antitumor CAR T-cell functionality in MM. This approach may also be effective for other cancers that depend on glutamine as a key energy source and metabolic hallmark.

Article Details

Journal Blood
Volume / Issue Vol. 146, Issue 24
Published December 11, 2025
Pages 2931-2944
ISSN 0006-4971
Publisher Elsevier BV

Journal Info

Blood

Elsevier BV

ISSN: 0006-4971 Health Sciences

Authors (26)

F

Flor Navarro

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

T

Teresa Lozano

Centro de Investigación Médica Aplicada, CIMA, Pamplona, Spain

A

Andrea Fuentes-García

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

I

Inés Sánchez-Moreno

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

M

Marta Larrayoz

P

Pedro Justicia

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

B

Beatriz Perucha

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

M

Maialen Martinez-Tabar

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

R

Rebeca Martinez-Turrillas

2Hemato-Oncology Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

N

Noelia Casares

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

C

Celia Martín-Otal

M

Marta Gorraiz

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

E

Erin W. Meermeier

1Department of Immunology, Mayo Clinic Arizona, Scottsdale, AZ

M

Marta Chesi

D

Douglas Lake

5School of Life Sciences, Arizona State University, Phoenix, AZ

P

P. Leif Bergsagel

Mayo Clinic Arizona, Scottsdale

E

Eva Santamaría

6RNA Biology and Therapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, University of Navarra, Pamplona, Spain

M

Maria Erendira Calleja-Cervantes

P

Patxi San Martín-Uriz

1Center for Applied Medical Research, Universidad de Navarra, Navarra Medical Research Institute, Pamplona, Spain

L

Lorea Jordana-Urriza

2Hemato-Oncology Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

X

Xabier Agirre

S

Sandra Hervás-Stubbs

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

J

Juan Roberto Rodriguez-Madoz

1Center for Applied Medical Research, Universidad de Navarra, Navarra Medical Research Institute, Pamplona, Spain

J

José Ángel Martinez-Climent

2Hemato-Oncology Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain

F

Felipe Prosper

J

Juan José Lasarte

1Immunology and Immunotherapy Program, Center for Applied Medical Research, Cancer Center University of Navarra, Navarra Institute for Health Research, CIBERONC, Pamplona, Spain