Redefining the Genetic Architecture of Hypertrophic Cardiomyopathy: Role of Intermediate-Effect Variants

S Soledad García Hernandez (Health in Code S.L, A Coruña, Spain (S.G.H.).) L Luis de la Higuera Romero (Health in Code S.L, A Coruña, Spain (L.d.l.H.R.).) A Adrian Fernandez M Maria Luisa Peña Peña (Department of Cardiology, Hospital Universitario Virgen del Rocío, Sevilla, Spain (M.L.P.P.).) N Nerea Mora-Ayestarán (Department of Cardiology, Hospital Universitario Puerta de Hierro, Instituto de Investigación Sanitaria Puerta de Hierro Majadahonda (IDIPHIM), Madrid, Spain (N.M.-A., N.R.-L., J.G.M., P.G.-P.).) M María Teresa Basurte-Elorz (Department of Cardiology, Área del Corazón, Hospital Universitario de Navarra, Pamplona, Spain (A.Y.I., M.S., G.L.-L., M.T.B.-E.).) J Jose María Larrañaga-Moreira (Unidad de Cardiopatías Familiares, Servicio de Cardiología, Complexo Hospitalario Universitario A Coruña, Instituto de Investigación Biomédica de A Coruña, Spain (J.M.L.-M., R.B.-V.).) I Ivonne Cárdenas Reyes (Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).) E Eduardo Villacorta (CIBER Cardiovascular, Instituto de Salud Carlos III, Madrid, Spain (C.G.-G., M.G.-D., G.C.-M., M.N.-P., J.M.G.-P., G.C.-V., A.B.-G., R.S.-B., A.G.-Á., M.B., E.Z., E.R.G., E.V., J.L.-F., M.S.-M., R.B.-V., A.D., M.A.E.-C., J.F.R.-P., J.G.M., P.G.-P.).) M Maria Valverde-Gómez (Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).) A Alicia Baustista-Paves (Inherited Cardiac Diseases Unit, San Cecilio Hospital. Granada, Spain (A.B.-P.).) E Elena Veira Villanueva (Complexo Hospitalario Universitario A Coruña, Spain (E.V.V.).) M Martín Ortiz-Genga (Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).) A Alex Lipov (Amsterdam UMC, University of Amsterdam, Amsterdam, Netherlands) N Noel Brogger (Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).) M María Sabater Molina (Inherited Cardiac Diseases Unit, Department of Cardiology, Hospital Universitario Virgen de la Arrixaca, El Palmar (Murcia), Spain (M.S.M.).) E Eduardo Moreno-Escobar (Cardiology Service, San Cecilio Clinical University Hospital, Granada, Spain (E.M.-E.).) L Luis Ruiz-Guerrero (Servicio de Cardiología, Hospital Universitario Marqués de Valdecilla, Santander, Cantabria, Spain (L.R.-G.).) P Petros Syrris (Institute of Cardiovascular Science, University College London, United Kingdom (A.P., I.H., P.S., P.M.E.).) X Xusto Fernández J Jesús Piqueras-Flores (Inherited Heart Disease and Heart Failure Unit, Cardiology Department, General University Hospital of Ciudad Real Faculty of Medicine, University of Castilla La Mancha IDISCAM, Biomedical Research Institute of Castilla La Manch, Spain (J.P.-F.).) A Almudena Amor Salamanca (Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).) C Connie R. Bezzina P Perry M. Elliott (Institute of Cardiovascular Science, University College London, United Kingdom (A.P., I.H., P.S., P.M.E.).) R Roberto Barriales-Villa (Complexo Hospitalario Universitario A Coruña, Instituto de Investigación Biomédica de A Coruña, CIBERCV–Instituto de Salud Carlos III, A Coruña, Spain) J Juan Ramón Gimeno-Blanes P Pablo Garcia-Pavia (Department of Cardiology, Hospital Universitario Puerta de Hierro, Instituto de Investigación Sanitaria Puerta de Hierro–Segovia de Arana, Centro de Investigación Biomédica en Red Enfermedades Cardiovaculares, and Centro Nacional de Investigaciones Cardiovasculares, Madrid) R Roddy Walsh (Cardiovascular and Genomics Research Institute, City St. George’s, University of London, London, UK (R.W.).) J Juan Pablo Ochoa (Centro Nacional de Investigaciones Cardiovasculares (CNIC), Madrid, Spain (N.R.-L., J.P.O., A.G.-Á., P.G.-P.).)

Abstract

BACKGROUND: Hypertrophic cardiomyopathy (HCM) is a genetically heterogeneous disorder linked primarily to rare variants in sarcomeric genes, although recently certain nonsarcomeric genes have emerged as important contributors. Nonmendelian genetic variants with reproducible moderate-effect sizes and low penetrance, intermediate-effect variants (IEVs), can play a crucial role in modulating disease expression. Understanding the clinical impact of IEVs is crucial to unravel the complex genetic architecture of HCM. METHODS: We conducted an ancestry-based enrichment analysis of 14 validated HCM genes, including the 9 core sarcomeric and 5 nonsarcomeric genes (ALPK3 , CSRP3 , FHOD3 , FLNC , and TRIM63 ). Enrichment of intermediate frequency missense variants was evaluated in 10 981 patients with HCM, 4030 internal controls of European-ancestry, and 590 000 external controls from gnomAD non-Finnish Europeans. The population-attributable fraction was calculated to assess contribution of IEVs to HCM. Age-related disease penetrance, phenotypic severity (left ventricular maximum wall thickness), and major adverse cardiac events were analyzed in 11 991 HCM cases of the whole cohort according to 5 genetic groups: genotype negative, isolated IEV, monogenic, monogenic+IEV, and double monogenic. RESULTS: Fourteen IEVs in 8 genes were identified in 731 individuals (6.1% of the cohort), of whom 570 patients (4.8%) had IEVs in isolation: 198 (34.7%) in sarcomeric genes and 372 (65.3%) in nonsarcomeric genes. The contribution of IEVs to HCM genetics according to population-attributable fraction was estimated to be 4.9% (95% CI, 3.2–6.7). A significant gradient in penetrance, phenotypic severity, and major adverse cardiac events was observed across genetic groups. Compared with genotype-negative patients, IEV carriers displayed a younger median age at diagnosis (59 years of age [95% CI, 46–69] versus 61 years [95% CI, 49–70]; P =0.0073) and a higher mean left ventricular maximum wall thickness (18.1±3.7 versus 19.0±4.3; P =0.0043). IEVs also modified disease expression in individuals with monogenic variants, causing a more aggressive phenotype than in individuals from the monogenic-only group with HCM onset at younger age and a higher left ventricular maximum wall thickness (all P <0.0001), with major adverse cardiac event–free survival being significantly lower (93.3% versus 69.3% at 70 years of age; P <0.0001). CONCLUSIONS: IEVs are present in 6.1% of HCM cases and account for 4.8% of HCM genetic burden. IEVs also influence disease severity and outcomes, particularly when combined with monogenic disease-causing variants. Evaluation of IEVs should be considered when HCM genetic testing is performed.

Article Details

Journal Circulation
Volume / Issue Vol. 152, Issue 15
Published October 14, 2025
Pages 1060-1075
ISSN 0009-7322
Publisher Lippincott Williams & Wilkins

Journal Info

Circulation

Lippincott Williams & Wilkins

ISSN: 0009-7322 Health Sciences

Authors (29)

S

Soledad García Hernandez

Health in Code S.L, A Coruña, Spain (S.G.H.).

L

Luis de la Higuera Romero

Health in Code S.L, A Coruña, Spain (L.d.l.H.R.).

A

Adrian Fernandez

M

Maria Luisa Peña Peña

Department of Cardiology, Hospital Universitario Virgen del Rocío, Sevilla, Spain (M.L.P.P.).

N

Nerea Mora-Ayestarán

Department of Cardiology, Hospital Universitario Puerta de Hierro, Instituto de Investigación Sanitaria Puerta de Hierro Majadahonda (IDIPHIM), Madrid, Spain (N.M.-A., N.R.-L., J.G.M., P.G.-P.).

M

María Teresa Basurte-Elorz

Department of Cardiology, Área del Corazón, Hospital Universitario de Navarra, Pamplona, Spain (A.Y.I., M.S., G.L.-L., M.T.B.-E.).

J

Jose María Larrañaga-Moreira

Unidad de Cardiopatías Familiares, Servicio de Cardiología, Complexo Hospitalario Universitario A Coruña, Instituto de Investigación Biomédica de A Coruña, Spain (J.M.L.-M., R.B.-V.).

I

Ivonne Cárdenas Reyes

Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).

E

Eduardo Villacorta

CIBER Cardiovascular, Instituto de Salud Carlos III, Madrid, Spain (C.G.-G., M.G.-D., G.C.-M., M.N.-P., J.M.G.-P., G.C.-V., A.B.-G., R.S.-B., A.G.-Á., M.B., E.Z., E.R.G., E.V., J.L.-F., M.S.-M., R.B.-V., A.D., M.A.E.-C., J.F.R.-P., J.G.M., P.G.-P.).

M

Maria Valverde-Gómez

Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).

A

Alicia Baustista-Paves

Inherited Cardiac Diseases Unit, San Cecilio Hospital. Granada, Spain (A.B.-P.).

E

Elena Veira Villanueva

Complexo Hospitalario Universitario A Coruña, Spain (E.V.V.).

M

Martín Ortiz-Genga

Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).

A

Alex Lipov

Amsterdam UMC, University of Amsterdam, Amsterdam, Netherlands

N

Noel Brogger

Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).

M

María Sabater Molina

Inherited Cardiac Diseases Unit, Department of Cardiology, Hospital Universitario Virgen de la Arrixaca, El Palmar (Murcia), Spain (M.S.M.).

E

Eduardo Moreno-Escobar

Cardiology Service, San Cecilio Clinical University Hospital, Granada, Spain (E.M.-E.).

L

Luis Ruiz-Guerrero

Servicio de Cardiología, Hospital Universitario Marqués de Valdecilla, Santander, Cantabria, Spain (L.R.-G.).

P

Petros Syrris

Institute of Cardiovascular Science, University College London, United Kingdom (A.P., I.H., P.S., P.M.E.).

X

Xusto Fernández

J

Jesús Piqueras-Flores

Inherited Heart Disease and Heart Failure Unit, Cardiology Department, General University Hospital of Ciudad Real Faculty of Medicine, University of Castilla La Mancha IDISCAM, Biomedical Research Institute of Castilla La Manch, Spain (J.P.-F.).

A

Almudena Amor Salamanca

Health in Code S.L., A Coruña, Spain (I.C.R., M.V.-G., M.O.-G., N.B., X.F., A.A.S., J.P.O.).

C

Connie R. Bezzina

P

Perry M. Elliott

Institute of Cardiovascular Science, University College London, United Kingdom (A.P., I.H., P.S., P.M.E.).

R

Roberto Barriales-Villa

Complexo Hospitalario Universitario A Coruña, Instituto de Investigación Biomédica de A Coruña, CIBERCV–Instituto de Salud Carlos III, A Coruña, Spain

J

Juan Ramón Gimeno-Blanes

P

Pablo Garcia-Pavia

Department of Cardiology, Hospital Universitario Puerta de Hierro, Instituto de Investigación Sanitaria Puerta de Hierro–Segovia de Arana, Centro de Investigación Biomédica en Red Enfermedades Cardiovaculares, and Centro Nacional de Investigaciones Cardiovasculares, Madrid

R

Roddy Walsh

Cardiovascular and Genomics Research Institute, City St. George’s, University of London, London, UK (R.W.).

J

Juan Pablo Ochoa

Centro Nacional de Investigaciones Cardiovasculares (CNIC), Madrid, Spain (N.R.-L., J.P.O., A.G.-Á., P.G.-P.).