Multimodal cell maps as a foundation for structural and functional genomics

L Leah V. Schaffer M Mengzhou Hu G Gege Qian K Kyung-Mee Moon A Abantika Pal N Neelesh Soni A Andrew P. Latham L Laura Pontano Vaites D Dorothy Tsai N Nicole M. Mattson K Katherine Licon R Robin Bachelder A Anthony Cesnik I Ishan Gaur T Trang Le W William Leineweber A Aji Palar E Ernst Pulido Y Yue Qin X Xiaoyu Zhao C Christopher Churas J Joanna Lenkiewicz J Jing Chen K Keiichiro Ono D Dexter Pratt P Peter Zage I Ignacia Echeverria A Andrej Sali J J. Wade Harper S Steven P. Gygi L Leonard J. Foster E Edward L. Huttlin E Emma Lundberg T Trey Ideker

Abstract

Abstract Human cells consist of a complex hierarchy of components, many of which remain unexplored1,2. Here we construct a global map of human subcellular architecture through joint measurement of biophysical interactions and immunofluorescence images for over 5,100 proteins in U2OS osteosarcoma cells. Self-supervised multimodal data integration resolves 275 molecular assemblies spanning the range of 10−8 to 10−5 m, which we validate systematically using whole-cell size-exclusion chromatography and annotate using large language models3. We explore key applications in structural biology, yielding structures for 111 heterodimeric complexes and an expanded Rag–Ragulator assembly. The map assigns unexpected functions to 975 proteins, including roles for C18orf21 in RNA processing and DPP9 in interferon signalling, and identifies assemblies with multiple localizations or cell type specificity. It decodes paediatric cancer genomes4, identifying 21 recurrently mutated assemblies and implicating 102 validated new cancer proteins. The associated Cell Visualization Portal and Mapping Toolkit provide a reference platform for structural and functional cell biology.

Article Details

Journal Nature
Volume / Issue Vol. 642, Issue 8066
Published June 05, 2025
Pages 222-231
ISSN 0028-0836
Publisher Nature Portfolio

Journal Info

Nature

Nature Portfolio

ISSN: 0028-0836 Health Sciences

Authors (34)

L

Leah V. Schaffer

M

Mengzhou Hu

G

Gege Qian

K

Kyung-Mee Moon

A

Abantika Pal

N

Neelesh Soni

A

Andrew P. Latham

L

Laura Pontano Vaites

D

Dorothy Tsai

N

Nicole M. Mattson

K

Katherine Licon

R

Robin Bachelder

A

Anthony Cesnik

I

Ishan Gaur

T

Trang Le

W

William Leineweber

A

Aji Palar

E

Ernst Pulido

Y

Yue Qin

X

Xiaoyu Zhao

C

Christopher Churas

J

Joanna Lenkiewicz

J

Jing Chen

K

Keiichiro Ono

D

Dexter Pratt

P

Peter Zage

I

Ignacia Echeverria

A

Andrej Sali

J

J. Wade Harper

S

Steven P. Gygi

L

Leonard J. Foster

E

Edward L. Huttlin

E

Emma Lundberg

T

Trey Ideker