Mitochondrial RNA cytosolic leakage drives the SASP

S Stella Victorelli M Madeline Eppard H Hélène Martini S Seung-Hwa Woo S Stacia P. A. Everts G Gung Lee N Nicholas Pirius N Nuan Han E Eugene Y. Liang A Ana Catarina Franco Y Yeaeun Han D Dominik Saul E Eva Nóvoa R Ruben Nogueiras P Patrick L. Splinter S Steven P. O’Hara O Olivia Morgenthaler L Lucía Valenzuela-Pérez H Hyun Se Kim Lee D Diana Jurk N Nicholas F. LaRusso P Petra Hirsova J João F. Passos

Abstract

Abstract Senescent cells secrete proinflammatory factors known as the senescence-associated secretory phenotype (SASP), contributing to tissue dysfunction and aging. Mitochondrial dysfunction is a key feature of senescence, influencing SASP via mitochondrial DNA (mtDNA) release and cGAS/STING pathway activation. Here, we demonstrate that mitochondrial RNA (mtRNA) also accumulates in the cytosol of senescent cells, activating RNA sensors RIG-I and MDA5, leading to MAVS aggregation and SASP induction. Inhibition of these RNA sensors significantly reduces SASP factors. Furthermore, BAX and BAK play a key role in mtRNA leakage during senescence, and their deletion diminishes SASP expression in vitro and in a mouse model of Metabolic Dysfunction-Associated Steatohepatitis (MASH). These findings highlight mtRNA’s role in SASP regulation and its potential as a therapeutic target for mitigating age-related inflammation.

Article Details

Volume / Issue Vol. 16, Issue 1
Published December 15, 2025
ISSN 2041-1723
Publisher Nature Portfolio

Journal Info

Nature Communications

Nature Portfolio

ISSN: 2041-1723 Open Access Life Sciences

Authors (23)

S

Stella Victorelli

M

Madeline Eppard

H

Hélène Martini

S

Seung-Hwa Woo

S

Stacia P. A. Everts

G

Gung Lee

N

Nicholas Pirius

N

Nuan Han

E

Eugene Y. Liang

A

Ana Catarina Franco

Y

Yeaeun Han

D

Dominik Saul

E

Eva Nóvoa

R

Ruben Nogueiras

P

Patrick L. Splinter

S

Steven P. O’Hara

O

Olivia Morgenthaler

L

Lucía Valenzuela-Pérez

H

Hyun Se Kim Lee

D

Diana Jurk

N

Nicholas F. LaRusso

P

Petra Hirsova

J

João F. Passos