Long-term cultural continuity across the Neanderthal–modern human sequence at Üçağızlı II Cave, northern Levant

İsmail Baykara (Department of Archaeology, Gaziantep University) D Didem Turan (Department of Archaeology, Karadeniz Technical University) E Ece Eren Kural (Department of Anthropology, Ankara University) D Derya Silibolatlaz (Department of Urbanization and Social Integration, Institute of Migration, Gaziantep University) M Mustafa Kenan Agras (Department of Archaeology, Gaziantep University) E Eren Şahiner (Earth Sciences Application and Research Center, Ankara University) S Salih Kavak (Department of Archaeology, Gaziantep University) C Clément Zanolli (De la Préhistoire à l’Actuel: Culture, Environnement et Anthropologie, Ministère de la Culture, UMR 5199, CNRS, University of Bordeaux) Y Yoshiro Ishihara (Department of Earth System Science, Faculty of Science, Fukuoka University) W Wataru Morita (Division of Anthropology, Department of Paleontology and Anthropology, National Museum of Nature and Science) N Naoki Morimoto (Laboratory of Physical Anthropology, Department of Zoology, Graduate School of Science, Kyoto University)

Abstract

We present evidence from Üçağızlı II Cave in the northern Levant, Türkiye, documenting the sequential occupation by Neanderthals ( Homo neanderthalensis ; 77–59 ka) and modern humans ( Homo sapiens ; 59–47 ka). The combined evidence of human fossils, faunal and floral remains, lithics, and manuports demonstrates significant behavioral and technological continuity across this taxonomic sequence, characterized by consistent subsistence strategies and the persistent selection and transport of specific nondietary mollusk shells. The lithic assemblages largely align with late Middle Paleolithic traditions, exhibiting characteristic Mousterian variants. The association of diagnostic human fossils with these archaeological remains demonstrates that the shift from Neanderthals to modern humans occurred within a cultural continuum. Our findings suggest shared behaviors between Neanderthals and modern humans that extended beyond subsistence to include nonutilitarian behaviors within the specific geographic and temporal context studied here.

Article Details

Volume / Issue Vol. 123, Issue 29
Published July 21, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (11)

İsmail Baykara

Department of Archaeology, Gaziantep University

D

Didem Turan

Department of Archaeology, Karadeniz Technical University

E

Ece Eren Kural

Department of Anthropology, Ankara University

D

Derya Silibolatlaz

Department of Urbanization and Social Integration, Institute of Migration, Gaziantep University

M

Mustafa Kenan Agras

Department of Archaeology, Gaziantep University

E

Eren Şahiner

Earth Sciences Application and Research Center, Ankara University

S

Salih Kavak

Department of Archaeology, Gaziantep University

C

Clément Zanolli

De la Préhistoire à l’Actuel: Culture, Environnement et Anthropologie, Ministère de la Culture, UMR 5199, CNRS, University of Bordeaux

Y

Yoshiro Ishihara

Department of Earth System Science, Faculty of Science, Fukuoka University

W

Wataru Morita

Division of Anthropology, Department of Paleontology and Anthropology, National Museum of Nature and Science

N

Naoki Morimoto

Laboratory of Physical Anthropology, Department of Zoology, Graduate School of Science, Kyoto University