Liver lipid droplet cholesterol content is a key determinant of metabolic dysfunction–associated steatohepatitis

I Ikki Sakuma (Department of Internal Medicine, Yale School of Medicine) R Rafael C. Gaspar (Department of Cellular & Molecular Physiology, Yale University) A Ali R. Nasiri (Department of Internal Medicine, Yale School of Medicine) S Sylvie Dufour (Department of Internal Medicine, Yale School of Medicine) M Mario Kahn (Department of Internal Medicine, Yale School of Medicine) J Jie Zheng (Key Laboratory of Radiation Physics and Technology, Ministry of Education, Institute of Nuclear Science and Technology) T Traci E. LaMoia (Department of Internal Medicine, Yale School of Medicine) M Mateus T. Guerra (Department of Internal Medicine, Yale School of Medicine) Y Yuki Taki (Department of Molecular Diagnosis, Chiba University Graduate School of Medicine) Y Yusuke Kawashima (Department of Applied Genomics, Kazusa deoxyribonucleic acid Research Institute) D Dean Yimlamai (Department of Pediatrics, Yale School of Medicine) M Mark Perelis (Ionis Pharmaceuticals) D Daniel F. Vatner (Department of Internal Medicine, Yale School of Medicine) K Kitt Falk Petersen (Department of Internal Medicine, Yale School of Medicine) M Maximilian Huttasch (Institute for Clinical Diabetology, German Diabetes Center (Deutsches Diabetes-Zentrum), Leibniz Center for Diabetes Research at Heinrich Heine University) B Birgit Knebel (German Center for Diabetes Research (Deutsche Zentrum für Diabetesforschung e.V.), Partner Düsseldorf) S Sabine Kahl (Institute for Clinical Diabetology, German Diabetes Center (Deutsches Diabetes-Zentrum), Leibniz Center for Diabetes Research at Heinrich Heine University) M Michael Roden (Institute for Clinical Diabetology, German Diabetes Center (Deutsches Diabetes-Zentrum), Leibniz Center for Diabetes Research at Heinrich Heine University) V Varman T. Samuel (Department of Internal Medicine, Yale School of Medicine) T Tomoaki Tanaka (Department of Molecular Diagnosis, Chiba University Graduate School of Medicine) G Gerald I. Shulman (Department of Internal Medicine, Yale School of Medicine)

Abstract

Metabolic dysfunction–associated steatohepatitis (MASH) represents a progressive form of steatotic liver disease which increases the risk for fibrosis and advanced liver disease. The accumulation of discrete species of bioactive lipids has been postulated to activate signaling pathways that promote inflammation and fibrosis. However, the key pathogenic lipid species is a matter of debate. We explored candidates using various dietary, molecular, and genetic models. Mice fed a choline-deficient L-amino acid–defined high-fat diet (CDAHFD) developed steatohepatitis and manifested early markers of liver fibrosis associated with increased cholesterol content in liver lipid droplets within 5 d without any changes in total liver cholesterol content. Treating mice with antisense oligonucleotides against Coenzyme A synthase ( Coasy ) or treatment with bempedoic acid or atorvastatin decreased liver lipid droplet cholesterol content and prevented CDAHFD-induced MASH and the fibrotic response. All these salutary effects were abrogated with dietary cholesterol supplementation. Analysis of human liver samples demonstrated that cholesterol in liver lipid droplets was increased in humans with MASH and liver fibrosis and was higher in PNPLA3 I148M (variants rs738409) than in HSD17B13 variants (rs72613567). Together, these data identify cholesterol in liver lipid droplets as a critical mediator of MASH and demonstrate that Coenzyme A synthase knockdown and bempedoic acid are therapeutic approaches to reduce liver lipid droplet cholesterol content and thereby prevent the development of MASH and liver fibrosis.

Article Details

Volume / Issue Vol. 122, Issue 18
Published May 06, 2025
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (21)

I

Ikki Sakuma

Department of Internal Medicine, Yale School of Medicine

R

Rafael C. Gaspar

Department of Cellular & Molecular Physiology, Yale University

A

Ali R. Nasiri

Department of Internal Medicine, Yale School of Medicine

S

Sylvie Dufour

Department of Internal Medicine, Yale School of Medicine

M

Mario Kahn

Department of Internal Medicine, Yale School of Medicine

J

Jie Zheng

Key Laboratory of Radiation Physics and Technology, Ministry of Education, Institute of Nuclear Science and Technology

T

Traci E. LaMoia

Department of Internal Medicine, Yale School of Medicine

M

Mateus T. Guerra

Department of Internal Medicine, Yale School of Medicine

Y

Yuki Taki

Department of Molecular Diagnosis, Chiba University Graduate School of Medicine

Y

Yusuke Kawashima

Department of Applied Genomics, Kazusa deoxyribonucleic acid Research Institute

D

Dean Yimlamai

Department of Pediatrics, Yale School of Medicine

M

Mark Perelis

Ionis Pharmaceuticals

D

Daniel F. Vatner

Department of Internal Medicine, Yale School of Medicine

K

Kitt Falk Petersen

Department of Internal Medicine, Yale School of Medicine

M

Maximilian Huttasch

Institute for Clinical Diabetology, German Diabetes Center (Deutsches Diabetes-Zentrum), Leibniz Center for Diabetes Research at Heinrich Heine University

B

Birgit Knebel

German Center for Diabetes Research (Deutsche Zentrum für Diabetesforschung e.V.), Partner Düsseldorf

S

Sabine Kahl

Institute for Clinical Diabetology, German Diabetes Center (Deutsches Diabetes-Zentrum), Leibniz Center for Diabetes Research at Heinrich Heine University

M

Michael Roden

Institute for Clinical Diabetology, German Diabetes Center (Deutsches Diabetes-Zentrum), Leibniz Center for Diabetes Research at Heinrich Heine University

V

Varman T. Samuel

Department of Internal Medicine, Yale School of Medicine

T

Tomoaki Tanaka

Department of Molecular Diagnosis, Chiba University Graduate School of Medicine

G

Gerald I. Shulman

Department of Internal Medicine, Yale School of Medicine