Interconnecting fragmented forests: Small and mobile birds are cornerstones in the plant–frugivore meta-network

C Chen Zhu B Bo Dalsgaard (Section for Molecular Ecology and Evolution, Globe Institute, University of Copenhagen) W Wande Li (Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University) C Christopher N. Kaiser-Bunbury (Centre for Ecology and Conservation, Faculty of Environment, Science and Economy, Penryn Campus, University of Exeter) B Benno I. Simmons (Centre for Ecology and Conservation, Faculty of Environment, Science and Economy, Penryn Campus, University of Exeter) P Peng Ren Y Yuhao Zhao (Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University) D Di Zeng (Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University) F Fernando Gonçalves (Section for Molecular Ecology and Evolution, Globe Institute, University of Copenhagen) X Xue Zhang L Longxiao Chang (Key Laboratory of Biosystems Homeostasis and Protection (Ministry of Education), College of Life Sciences, Zhejiang University) P Ping Ding (State Key Laboratory of Deep Earth Processes and Resources, Guangzhou Institute of Geochemistry, Chinese Academy of Sciences) X Xingfeng Si (Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University)

Abstract

Habitat fragmentation is causing the collapse of seed dispersal interactions and ecosystem functioning. When management and conservation strategies aim to sustain ecosystem functioning of fragmented forests, species’ traits and functional performance are critical in guiding decisions. However, to date, we lack a quantitative understanding of the role of frugivores’ body size and dispersal ability in ecosystem sustainability among fragmented forests. Focusing on avian frugivory and seed dispersal in a multi-island setting, we address the data gap by recording more than 20,000 frugivory events in an artificial insular fragmented landscape constructed in 1959 and nearby unfragmented forests on the mainland. We show that large-bodied and dispersal-limited frugivorous birds are largely confined to large islands and the unfragmented mainland, whereas on small islands, small-bodied and highly mobile birds predominantly engage in frugivory interactions. The plant–frugivore meta-network exhibits a distinct compartmentalization, driven by island area and bird mobility. Birds with smaller size and greater mobility have higher topological importance, and the presence of small-bodied birds significantly enhances meta-network robustness. These results suggest that among insular fragmented forests where frugivory interactions are degraded, small-bodied and highly mobile birds disproportionately contribute to meta-community cohesion and ecosystem functioning because of the lack of large-bodied and dispersal-limited birds. We thus advocate for the restoration of landscapes to facilitate seed dispersal and functional connectivity, ensuring the presence of large patches along with small patches as stepping-stones. Meanwhile, we recommend prioritizing conservation on small-bodied and highly mobile birds in fragmented landscapes, a subset of underappreciated species that yet play crucial roles in ecosystem functioning.

Article Details

Volume / Issue Vol. 122, Issue 7
Published February 18, 2025
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (13)

C

Chen Zhu

B

Bo Dalsgaard

Section for Molecular Ecology and Evolution, Globe Institute, University of Copenhagen

W

Wande Li

Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University

C

Christopher N. Kaiser-Bunbury

Centre for Ecology and Conservation, Faculty of Environment, Science and Economy, Penryn Campus, University of Exeter

B

Benno I. Simmons

Centre for Ecology and Conservation, Faculty of Environment, Science and Economy, Penryn Campus, University of Exeter

P

Peng Ren

Y

Yuhao Zhao

Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University

D

Di Zeng

Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University

F

Fernando Gonçalves

Section for Molecular Ecology and Evolution, Globe Institute, University of Copenhagen

X

Xue Zhang

L

Longxiao Chang

Key Laboratory of Biosystems Homeostasis and Protection (Ministry of Education), College of Life Sciences, Zhejiang University

P

Ping Ding

State Key Laboratory of Deep Earth Processes and Resources, Guangzhou Institute of Geochemistry, Chinese Academy of Sciences

X

Xingfeng Si

Center for Global Change and Ecological Forecasting, Zhejiang Zhoushan Island Ecosystem Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University