Infra-slow (<0.1 Hz) Modulation of Human Brain Pulsations in Awake and Sleep States

T Tommi Väyrynen (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) H Heta Helakari (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) V Vesa Korhonen (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) J Johanna Tuunanen (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) N Niko Huotari (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) J Janne Kananen (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) S Seyed-Mohsen Ebrahimi A Ahmed Elabasy (Oulu Functional NeuroImaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) M Matti Järvelä (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) K Katariina Hautamäki (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) K Katariina Laurén L Lauri Raitamaa (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) U Ulla Salmi J Johanna Piispala (Clinical Neurophysiology, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) M Mika Kallio (Clinical Neurophysiology, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu) V Vesa Kiviniemi (Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu)

Abstract

The human brain exhibits three propagating pulsations, namely, cardiovascular, respiratory, and infra-slow fluctuations (ISFs <0.1 Hz), which are thought to contribute to the flow of intracranial fluids. While their pulsation characteristics have been extensively studied, their mutual dependencies have not been systematically investigated. Using ultrafast whole-brain magnetic resonance encephalography fMRI sequence, we analyzed the frequency domain up to 5 Hz for cross-frequency oscillatory interactions in awake and NREM sleep states of 23 (13 F, 10 M) healthy volunteers. Using phase transfer entropy analysis, we found that in the awake state the resting-state network (RSN) activity largely predicted the neurofluid (NF) signal changes. NREM sleep was associated with increased power of ISF and with altered directed coupling patterns between RSN and NF components. Importantly, within these independent signal sources, we found three distinct cross-frequency coupling frequency ranges occurring at ISF (<0.1 Hz), respiratory (∼0.25 Hz), and cardiovascular (∼1 Hz) frequencies, where the slower pulsations generally predicted the faster ones, except for a finding of inverted cardiorespiratory coupling in NREM-2 sleep. These results suggest the presence of directional ISF mediated mechanisms underlying brain pulsations that contribute to driving the intracranial fluid transfer processes.

Article Details

Volume / Issue Vol. 46, Issue 27
Published July 08, 2026
Pages e2282252026
ISSN 0270-6474
Publisher Society for Neuroscience

Journal Info

Journal of Neuroscience

Society for Neuroscience

ISSN: 0270-6474 Life Sciences

Authors (16)

T

Tommi Väyrynen

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

H

Heta Helakari

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

V

Vesa Korhonen

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

J

Johanna Tuunanen

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

N

Niko Huotari

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

J

Janne Kananen

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

S

Seyed-Mohsen Ebrahimi

A

Ahmed Elabasy

Oulu Functional NeuroImaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

M

Matti Järvelä

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

K

Katariina Hautamäki

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

K

Katariina Laurén

L

Lauri Raitamaa

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

U

Ulla Salmi

J

Johanna Piispala

Clinical Neurophysiology, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

M

Mika Kallio

Clinical Neurophysiology, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu

V

Vesa Kiviniemi

Oulu Functional Neuroimaging, Research unit of Health Sciences and Technology, Faculty of Medicine, Medical Research Center, University of Oulu