IL-16 production is a mechanism of resistance to BTK inhibitors and R-CHOP in lymphomas

A Alberto J Arribas (Università della Svizzera italiana, Institute of Oncology Research, Bellinzona, Switzerland) E Eleonora Cannas (1 Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland, Bellinzona, Switzerland) G Giulio Sartori (Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland) F Francesca Guidetti (Università della Svizzera italiana, Istituto Oncologico di Ricerca) L Luciano Cascione (IOR - Institute of Oncology Research, Bellinzona, Switzerland) S Sara Napoli (Institute of Oncology Research, Bellinzona, Switzerland) G Gabriela Forestieri (Institute of Oncology Research, Bellinzona, Switzerland) F Federica Fuzio (Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland) E Emma Daniela Susanna Pesenti (Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland) C Chiara Tarantelli (Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland) F Filippo Spriano (Institute of Oncology Research, Faculty of Biomedical Sciences, USI, bellinzona, Switzerland) A Antonella Zucchetto (3Clinical and Experimental Onco-Hematology Unit, Centro di Riferimento Oncologico di Aviano, Istituto di Ricovero e Cura a Carattere Scientifico, Aviano, Italy) F Francesca Maria Rossi (Centro di Riferimento Oncologico di Aviano (CRO) IRCCS, Aviano, Italy) A Alessio Bruscaggin (Institute of Oncology Research, Bellinzona, Switzerland) A Andrea Rinaldi M Manuel Castro de Moura (Josep Carreras Leukaemia Research Institute, Badalona, Spain) S Sandra Jovic (Institute for Research in Biomedicine, Università della Svizzera italiana, Bellinzona, Switzerland) A Andrea Raimondi (Protein Folding and Quality Control, Faculty of Biomedical Sciences, Institute for Research in Biomedicine, Università della Svizzera italiana) R Roberta Bordone Pittau (IOSI Oncology Institute of Southern Switzerland) L Lodovico Terzi di Bergamo (Institute of Oncology Research, Bellinzona, Switzerland) X Xiaofei Ye A Anastasios Stathis (Oncology Institute of Southern Switzerland, Ente Ospedaliero Cantonale, Bellinzoona, Switzerland) Y Yaacov Ben-David (State Key Laboratory for Functions and Applications of Medicinal Plants, Guiyang, China) Q Qiang Pan-Hammarström (Division of Immunology, Department of Medical Biochemistry and Biophysics, Karolinska Institute) F Federico Simonetta (Geneva University Hospitals and University of Geneva, Genève, Switzerland) G Georg Stussi (Oncology Institute of Southern Switzerland, Bellinzona, Switzerland) E Emanuele Zucca (Institute of Oncology Research, Bellinzona, Switzerland) V Valter Gattei (3Clinical and Experimental Onco-Hematology Unit, Centro di Riferimento Oncologico di Aviano, Istituto di Ricovero e Cura a Carattere Scientifico, Aviano, Italy) J Jennifer R. Brown M Manel Esteller D Davide Rossi (Institute of Oncology Research, Bellinzona, Switzerland) F Francesco Bertoni (Oncology Institute of Southern Switzerland, Ente Ospedaliero Cantonale, Bellinzona, Switzerland)

Abstract

Resistance to Bruton tyrosine kinase (BTK) inhibitors remains a major clinical challenge in B-cell lymphomas and often occurs in the absence of BTK or PLCG2 mutations. Here, we investigated non-genetic mechanisms of ibrutinib resistance in marginal zone lymphoma (MZL) and their broader therapeutic implications. Chronic ibrutinib exposure generated a resistant MZL model that showed cross-resistance to BTK inhibitors and degraders, without evidence of multidrug resistance or genetic alterations. Integrated transcriptomic, epigenetic, and proteomic analyses revealed extensive reprogramming, including activation of PI3K/AKT, MAPK, and MYC pathways, repression of apoptosis and oxidative phosphorylation, and a prominent cytokine-secretory phenotype. Interleukin-16 (IL-16) emerged as a central mediator of resistance. IL-16 was transcriptionally upregulated, actively secreted, and sufficient to induce ibrutinib resistance across multiple CD9-positive models of MZL, mantle cell lymphoma, chronic lymphocytic leukemia (CLL), and activated B-cell-like diffuse large B-cell lymphoma. Serum IL-16 levels were elevated in ibrutinib-refractory CLL patients without BTK/PLCG2 mutations. Mechanistically, IL-16 engaged CD9-enriched membrane microdomains to activate PI3Kδ, thereby sustaining AKT and ERK signaling, stabilizing MYC, inducing NF-κB-dependent programs, and upregulating antiapoptotic effectors, including BFL1 (BCL2A1). Pharmacological or genetic disruption of the IL-16/CD9/PI3K axis restored sensitivity to BTK inhibitors and R-CHOP and abrogated IL-16-induced signaling in primary CLL samples. In conclusion, an IL-16/CD9-driven, epigenetically regulated survival pathway represents one possible mechanism of resistance to BTK inhibitors and chemoimmunotherapy, supporting therapeutic targeting of this axis in refractory B-cell lymphomas.

Article Details

Journal Blood
Volume / Issue Vol. 1, Issue 1
Published May 29, 2026
ISSN 0006-4971
Publisher Elsevier BV

Journal Info

Blood

Elsevier BV

ISSN: 0006-4971 Health Sciences

Authors (32)

A

Alberto J Arribas

Università della Svizzera italiana, Institute of Oncology Research, Bellinzona, Switzerland

E

Eleonora Cannas

1 Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland, Bellinzona, Switzerland

G

Giulio Sartori

Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland

F

Francesca Guidetti

Università della Svizzera italiana, Istituto Oncologico di Ricerca

L

Luciano Cascione

IOR - Institute of Oncology Research, Bellinzona, Switzerland

S

Sara Napoli

Institute of Oncology Research, Bellinzona, Switzerland

G

Gabriela Forestieri

Institute of Oncology Research, Bellinzona, Switzerland

F

Federica Fuzio

Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland

E

Emma Daniela Susanna Pesenti

Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland

C

Chiara Tarantelli

Institute of Oncology Research, Faculty of Biomedical Sciences, USI, Bellinzona, Switzerland

F

Filippo Spriano

Institute of Oncology Research, Faculty of Biomedical Sciences, USI, bellinzona, Switzerland

A

Antonella Zucchetto

3Clinical and Experimental Onco-Hematology Unit, Centro di Riferimento Oncologico di Aviano, Istituto di Ricovero e Cura a Carattere Scientifico, Aviano, Italy

F

Francesca Maria Rossi

Centro di Riferimento Oncologico di Aviano (CRO) IRCCS, Aviano, Italy

A

Alessio Bruscaggin

Institute of Oncology Research, Bellinzona, Switzerland

A

Andrea Rinaldi

M

Manuel Castro de Moura

Josep Carreras Leukaemia Research Institute, Badalona, Spain

S

Sandra Jovic

Institute for Research in Biomedicine, Università della Svizzera italiana, Bellinzona, Switzerland

A

Andrea Raimondi

Protein Folding and Quality Control, Faculty of Biomedical Sciences, Institute for Research in Biomedicine, Università della Svizzera italiana

R

Roberta Bordone Pittau

IOSI Oncology Institute of Southern Switzerland

L

Lodovico Terzi di Bergamo

Institute of Oncology Research, Bellinzona, Switzerland

X

Xiaofei Ye

A

Anastasios Stathis

Oncology Institute of Southern Switzerland, Ente Ospedaliero Cantonale, Bellinzoona, Switzerland

Y

Yaacov Ben-David

State Key Laboratory for Functions and Applications of Medicinal Plants, Guiyang, China

Q

Qiang Pan-Hammarström

Division of Immunology, Department of Medical Biochemistry and Biophysics, Karolinska Institute

F

Federico Simonetta

Geneva University Hospitals and University of Geneva, Genève, Switzerland

G

Georg Stussi

Oncology Institute of Southern Switzerland, Bellinzona, Switzerland

E

Emanuele Zucca

Institute of Oncology Research, Bellinzona, Switzerland

V

Valter Gattei

3Clinical and Experimental Onco-Hematology Unit, Centro di Riferimento Oncologico di Aviano, Istituto di Ricovero e Cura a Carattere Scientifico, Aviano, Italy

J

Jennifer R. Brown

M

Manel Esteller

D

Davide Rossi

Institute of Oncology Research, Bellinzona, Switzerland

F

Francesco Bertoni

Oncology Institute of Southern Switzerland, Ente Ospedaliero Cantonale, Bellinzona, Switzerland