<i>BCL11B</i> enhancer hijacking by t(14;16)(q32;q24) translocation defines a novel high-risk subtype of T-ALL

K Kaito Mimura (1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan) A Akira Kaino (1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan) Y Yotaro Ochi Y Yu-Hsuan Chang (4Division of Molecular Oncology, Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan) M Masafumi Seki J June Takeda (1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan) S Saori Katayama (2Department of Pediatrics, Tohoku University Graduate School of Medicine, Sendai, Japan) H Hidetaka Niizuma Y Yoji Sasahara (Department of Pediatrics, Tohoku University Graduate School of Medicine) Y Yoko Mizoguchi (7Department of Pediatrics, Hiroshima University Hospital, Hiroshima, Japan) M Maiko Shimomura (7Department of Pediatrics, Hiroshima University Hospital, Hiroshima, Japan) R Ryosuke Koyamada (8Department of Hematology, St. Luke’s International Hospital, Tokyo, Japan) R Rintaro Ono (9Department of Pediatrics, St. Luke's International Hospital, Tokyo, Japan) D Daisuke Hasegawa (11Department of Pediatrics, St. Luke's International Hospital, Tokyo, Japan) K Kazuki Mitani (10Department of Pediatrics, Graduate School of Medicine, Kyoto University, Kyoto, Japan) H Hirohito Kubota S Satoshi Yoshihara N Nobuhiro Hiramoto A Akihito Otsuki Y Yasunobu Okamura F Fumiki Katsuoka K Kengo Kinoshita M Masataka Hasegawa (1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan) M Marina Togo-Ohno (1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan) H Hirona Maeda (3Department of Pathology and Tumor Biology, Graduate School of Medicine, Kyoto University, Kyoto, Japan) N Nobuyuki Kakiuchi M Mai Takeuchi (19Department of Diagnostic Pathology, National Cancer Center Hospital, Tokyo, Japan) A Aiko Sato-Otsubo (6Department of Pediatrics, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan) S Shota Kato K Kentaro Watanabe (6Department of Pediatrics, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan) K Kotoe Katayama S Seiya Imoto Y Yuichi Shiraishi K Katsuyoshi Koh (23Japan Children’s Cancer Group ALL Committee, Nagoya, Japan) S Souichi Suenobu (25Japan Association of Childhood Leukemia Study, Kobe, Japan) E Eiso Hiyama (27Department of Biomedical Science, Natural Science Center for Basic Research and Development, Hiroshima University, Hiroshima, Japan) S Susumu Goyama (4Division of Molecular Oncology, Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan) A Atsuo Kikuchi S Seishi Ogawa M Motohiro Kato (6Department of Pediatrics, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan) Y Yasuhito Nannya J Junko Takita K Kenichi Yoshida

Abstract

Abstract The molecular classification of T-cell acute lymphoblastic leukemia (T-ALL) remains incomplete, limiting risk stratification and the development of targeted therapies. Enhancer hijacking is a critical oncogenic mechanism that deregulates proto-oncogenes by repositioning cisregulatory regions via structural variants. Here, we performed an integrated analysis of pediatric and adult T-ALL and mixed-phenotype acute leukemias (MPALs), using whole-genome and whole-transcriptome sequencing. This analysis identified a group of 14 patients with predominantly T-lineage neoplasms driven by a t(14;16)(q32;q24) translocation, harboring universal GATA3 mutations and CDKN2A/B deletions. Mechanistically, this translocation repositions the ThymoD locus downstream of BCL11B, causing monoallelic, ectopic overexpression of FENDRR and mesenchymal transcription factor genes FOXF1 and FOXC2 and activating epithelial-mesenchymal transition transcription signatures. Immunophenotypic and single-cell RNA sequencing analyses revealed marked lineage ambiguity with myeloid and B-cell differentiation potentials specific to this subtype. Furthermore, functional analyses in CD34+ cord blood cells demonstrated that FOXF1 overexpression promotes myeloid differentiation while suppressing T-cell differentiation, serving as a key factor for lineage specification. Clinically, this subtype was detected in 0.15% to 4.0% of T-ALL/MPAL cases depending on the cohort, showing a median age of 15 years and enrichment in adolescents and young adults. Importantly, patients with t(14;16)(q32;q24) have an extremely poor prognosis, showing a trend toward worse outcomes than high-risk groups such as KMT2A-rearranged early T-cell progenitor-like, SPI1-rearranged, and LMO2 γδ-like T-ALLs. The unique molecular landscape and poor prognosis of patients with the t(14;16)(q32;q24) translocation underscore the need for the development of novel subtype-specific therapeutic approaches.

Article Details

Journal Blood
Volume / Issue Vol. 148, Issue 3
Published July 16, 2026
Pages 335-347
ISSN 0006-4971
Publisher Elsevier BV

Journal Info

Blood

Elsevier BV

ISSN: 0006-4971 Health Sciences

Authors (43)

K

Kaito Mimura

1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan

A

Akira Kaino

1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan

Y

Yotaro Ochi

Y

Yu-Hsuan Chang

4Division of Molecular Oncology, Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan

M

Masafumi Seki

J

June Takeda

1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan

S

Saori Katayama

2Department of Pediatrics, Tohoku University Graduate School of Medicine, Sendai, Japan

H

Hidetaka Niizuma

Y

Yoji Sasahara

Department of Pediatrics, Tohoku University Graduate School of Medicine

Y

Yoko Mizoguchi

7Department of Pediatrics, Hiroshima University Hospital, Hiroshima, Japan

M

Maiko Shimomura

7Department of Pediatrics, Hiroshima University Hospital, Hiroshima, Japan

R

Ryosuke Koyamada

8Department of Hematology, St. Luke’s International Hospital, Tokyo, Japan

R

Rintaro Ono

9Department of Pediatrics, St. Luke's International Hospital, Tokyo, Japan

D

Daisuke Hasegawa

11Department of Pediatrics, St. Luke's International Hospital, Tokyo, Japan

K

Kazuki Mitani

10Department of Pediatrics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

H

Hirohito Kubota

S

Satoshi Yoshihara

N

Nobuhiro Hiramoto

A

Akihito Otsuki

Y

Yasunobu Okamura

F

Fumiki Katsuoka

K

Kengo Kinoshita

M

Masataka Hasegawa

1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan

M

Marina Togo-Ohno

1Division of Cancer Evolution, National Cancer Center Research Institute, Tokyo, Japan

H

Hirona Maeda

3Department of Pathology and Tumor Biology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

N

Nobuyuki Kakiuchi

M

Mai Takeuchi

19Department of Diagnostic Pathology, National Cancer Center Hospital, Tokyo, Japan

A

Aiko Sato-Otsubo

6Department of Pediatrics, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

S

Shota Kato

K

Kentaro Watanabe

6Department of Pediatrics, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

K

Kotoe Katayama

S

Seiya Imoto

Y

Yuichi Shiraishi

K

Katsuyoshi Koh

23Japan Children’s Cancer Group ALL Committee, Nagoya, Japan

S

Souichi Suenobu

25Japan Association of Childhood Leukemia Study, Kobe, Japan

E

Eiso Hiyama

27Department of Biomedical Science, Natural Science Center for Basic Research and Development, Hiroshima University, Hiroshima, Japan

S

Susumu Goyama

4Division of Molecular Oncology, Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan

A

Atsuo Kikuchi

S

Seishi Ogawa

M

Motohiro Kato

6Department of Pediatrics, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Y

Yasuhito Nannya

J

Junko Takita

K

Kenichi Yoshida