High-quality surrounding landscapes mitigate avian extirpations from forest remnants

A Anderson S. Bueno (Instituto Federal de Educação, Ciência e Tecnologia Farroupilha) C Chase D. Mendenhall (Physician Assistant Studies, Slippery Rock University) M Marina Anciães (Instituto Nacional de Pesquisas da Amazônia) L Luiz dos Anjos (Departamento de Biologia Animal e Vegetal, Universidade Estadual de Londrina) V Víctor Arroyo-Rodríguez (Instituto de Investigaciones en Ecosistemas y Sustentabilidad, Universidad Nacional Autónoma de México) M Marco Aurélio-Silva (Instituto Nacional de Pesquisas da Amazônia) C Cristina Banks-Leite (Department of Life Sciences, Imperial College London) M Matthew G. Betts (Department of Forest Ecosystems and Society, Oregon State University) A Arthur A. Bispo (Laboratório de Etnobiologia e Biodiversidade, Universidade Federal de Goiás) A Andrea Larissa Boesing (Senckenberg Biodiversity and Climate Research Centre, Senckenberg Gesellschaft für Naturforschung) M Marconi Campos-Cerqueira (WildMon) O Olivier Claessens (Groupe d’Étude et de Protection des Oiseaux en Guyane) W Will Cresswell (Centre for Biological Diversity, University of St. Andrews) G Gretchen C. Daily (Department of Biology, Stanford University) F Filibus Danjuma Dami (A.P. Leventis Ornithological Research Institute, University of Jos) S Sidnei M. Dantas (Instituto Tecnológico Vale) P Pedro F. Develey (SAVE Brasil) P Ping Ding (State Key Laboratory of Deep Earth Processes and Resources, Guangzhou Institute of Geochemistry, Chinese Academy of Sciences) D David P. Edwards M Márcio A. Efe (Laboratório de Bioecologia e Conservação de Aves Neotropicais, Universidade Federal de Alagoas) D Deborah Faria K Kenneth J. Feeley (Department of Biology, University of Miami) T Thomas W. Gillespie (Department of Geography, University of California) A Adam S. Hadley (New Brunswick Department of Natural Resources and Energy Development) J Jack H. Hatfield (Leverhulme Centre for Anthropocene Biodiversity and Department of Biology, University of York) L Luiza Magalli Pinto Henriques (Instituto Nacional de Pesquisas da Amazônia) L Lars H. Holbech (Department of Animal Biology and Conservation Science, University of Ghana) G Gregory J. Irving (Conservation Ecology Program, King Mongkut’s University of Technology Thonburi) U Urs G. Kormann (Swiss Ornithological Institute) M M. Jyothi Krishnan (Department of Wildlife Science, Kerala Agricultural University) M Marilise M. Krügel (Departamento de Engenharia Sanitária e Ambiental, Universidade Federal de Santa Maria) J Jesse R. Lasky M Michael J. Lawes (School of Life Sciences, University of KwaZulu-Natal) A Alexander C. Lees L Luc Lens (Centre for Research on Ecology, Cognition and Behaviour of Birds, Ghent University) L Lahert William Lobo-Araújo (Programa de Pós-Graduação em Ecologia e Recursos Naturais, Universidade Federal de São Carlos) L Leithen K. M’Gonigle (Department of Biological Sciences, Simon Fraser University) M Mohammad Saiful Mansor (Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, Universiti Kebangsaan Malaysia) S Shiiwua A. Manu (A.P. Leventis Ornithological Research Institute, University of Jos) M Miguel Ângelo Marini (Departamento de Zoologia, Universidade de Brasília) A Alexandre Camargo Martensen (Centro de Ciências da Natureza, Universidade Federal de São Carlos) T Thomas J. Matthews (School of Geography, Earth and Environmental Sciences and Birmingham Institute of Forest Research, University of Birmingham) J Jean Paul Metzger R Randall Moore (Department of Fisheries, Wildlife, and Conservation Sciences, Oregon State University) J José Carlos Morante-Filho (Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz) P P. O. Nameer (Department of Wildlife Science, Kerala Agricultural University) S Shukor Md Nor (Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, Universiti Kebangsaan Malaysia) H Helon Simões Oliveira (Programa de Pós-Graduação em Ecologia e Conservação, Universidade Federal de Sergipe) R Rômulo Ribon (Museu de Zoologia João Moojen, Departamento de Biologia Animal, Universidade Federal de Viçosa) V Viviana Ruiz-Gutiérrez (Cornell Lab of Ornithology, Cornell University) L Luís Fábio Silveira (Museu de Zoologia da Universidade de São Paulo) P Philip C Stouffer (School of Renewable Natural Resources, Louisiana State University AgCenter and Louisiana State University) J John W. Terborgh (Department of Biology, University of Florida) A Alexandre Uezu (Instituto de Pesquisas Ecológicas) Y Yanping Wang (Zhejiang Engineering Research Center for Biological Control of Crop Pathogens and Insects, Zhejiang Key Laboratory of Biology and Ecological Regulation of Crop Pathogens and Insects, Ministry of Agriculture and Rural Affairs Key Lab of Molecular Biology of Crop Pathogens and Insect Pests, State Key Lab of Rice Biology and Breeding, Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University) R Robyn Wethered (Cossypha Ecological) D Ding Li Yong (BirdLife International (Asia), Tanglin International Centre) C Carlos A. Peres

Abstract

The species–area relationship (SAR) has long been used to predict extirpation rates from habitat loss, but these rates depend not only on habitat area but also on the surrounding landscape and species’ habitat specialization. We collated global data from forest islands created by river damming and forest fragments resulting from clear-cut deforestation to examine the effects of matrix type (aquatic or terrestrial) and tree cover on avian SARs. Unlike oceanic islands, which are often millions of years old, anthropogenic forest islands provide a contemporary analog to forest fragments to understand matrix effects on SARs and serve as a baseline for worst-case scenarios of forest fragmentation. Our database comprises 50 datasets from 45 studies conducted in tropical and subtropical regions, totaling 1,954 bird species detected through 39,197 incidence records from 336 forest islands and 669 forest fragments. We found that bird extirpation rates were lower in fragments than on islands, especially for forest-dependent species compared to all species. Species losses were further reduced by increasing tree cover around forest remnants at local landscape scales of 300 m, highlighting the importance of small-scale conservation strategies. Moreover, even small forest fragments with greater nearby tree cover held high conservation value, emphasizing the crucial role of the surrounding landscape in mitigating avian extirpations from forest remnants. Beyond protecting forest remnants themselves, area-based conservation efforts would therefore be greatly enhanced by improving matrix quality and expanding tree cover in otherwise hostile landscapes.

Article Details

Volume / Issue Vol. 123, Issue 14
Published April 07, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (58)

A

Anderson S. Bueno

Instituto Federal de Educação, Ciência e Tecnologia Farroupilha

C

Chase D. Mendenhall

Physician Assistant Studies, Slippery Rock University

M

Marina Anciães

Instituto Nacional de Pesquisas da Amazônia

L

Luiz dos Anjos

Departamento de Biologia Animal e Vegetal, Universidade Estadual de Londrina

V

Víctor Arroyo-Rodríguez

Instituto de Investigaciones en Ecosistemas y Sustentabilidad, Universidad Nacional Autónoma de México

M

Marco Aurélio-Silva

Instituto Nacional de Pesquisas da Amazônia

C

Cristina Banks-Leite

Department of Life Sciences, Imperial College London

M

Matthew G. Betts

Department of Forest Ecosystems and Society, Oregon State University

A

Arthur A. Bispo

Laboratório de Etnobiologia e Biodiversidade, Universidade Federal de Goiás

A

Andrea Larissa Boesing

Senckenberg Biodiversity and Climate Research Centre, Senckenberg Gesellschaft für Naturforschung

M

Marconi Campos-Cerqueira

WildMon

O

Olivier Claessens

Groupe d’Étude et de Protection des Oiseaux en Guyane

W

Will Cresswell

Centre for Biological Diversity, University of St. Andrews

G

Gretchen C. Daily

Department of Biology, Stanford University

F

Filibus Danjuma Dami

A.P. Leventis Ornithological Research Institute, University of Jos

S

Sidnei M. Dantas

Instituto Tecnológico Vale

P

Pedro F. Develey

SAVE Brasil

P

Ping Ding

State Key Laboratory of Deep Earth Processes and Resources, Guangzhou Institute of Geochemistry, Chinese Academy of Sciences

D

David P. Edwards

M

Márcio A. Efe

Laboratório de Bioecologia e Conservação de Aves Neotropicais, Universidade Federal de Alagoas

D

Deborah Faria

K

Kenneth J. Feeley

Department of Biology, University of Miami

T

Thomas W. Gillespie

Department of Geography, University of California

A

Adam S. Hadley

New Brunswick Department of Natural Resources and Energy Development

J

Jack H. Hatfield

Leverhulme Centre for Anthropocene Biodiversity and Department of Biology, University of York

L

Luiza Magalli Pinto Henriques

Instituto Nacional de Pesquisas da Amazônia

L

Lars H. Holbech

Department of Animal Biology and Conservation Science, University of Ghana

G

Gregory J. Irving

Conservation Ecology Program, King Mongkut’s University of Technology Thonburi

U

Urs G. Kormann

Swiss Ornithological Institute

M

M. Jyothi Krishnan

Department of Wildlife Science, Kerala Agricultural University

M

Marilise M. Krügel

Departamento de Engenharia Sanitária e Ambiental, Universidade Federal de Santa Maria

J

Jesse R. Lasky

M

Michael J. Lawes

School of Life Sciences, University of KwaZulu-Natal

A

Alexander C. Lees

L

Luc Lens

Centre for Research on Ecology, Cognition and Behaviour of Birds, Ghent University

L

Lahert William Lobo-Araújo

Programa de Pós-Graduação em Ecologia e Recursos Naturais, Universidade Federal de São Carlos

L

Leithen K. M’Gonigle

Department of Biological Sciences, Simon Fraser University

M

Mohammad Saiful Mansor

Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, Universiti Kebangsaan Malaysia

S

Shiiwua A. Manu

A.P. Leventis Ornithological Research Institute, University of Jos

M

Miguel Ângelo Marini

Departamento de Zoologia, Universidade de Brasília

A

Alexandre Camargo Martensen

Centro de Ciências da Natureza, Universidade Federal de São Carlos

T

Thomas J. Matthews

School of Geography, Earth and Environmental Sciences and Birmingham Institute of Forest Research, University of Birmingham

J

Jean Paul Metzger

R

Randall Moore

Department of Fisheries, Wildlife, and Conservation Sciences, Oregon State University

J

José Carlos Morante-Filho

Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz

P

P. O. Nameer

Department of Wildlife Science, Kerala Agricultural University

S

Shukor Md Nor

Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, Universiti Kebangsaan Malaysia

H

Helon Simões Oliveira

Programa de Pós-Graduação em Ecologia e Conservação, Universidade Federal de Sergipe

R

Rômulo Ribon

Museu de Zoologia João Moojen, Departamento de Biologia Animal, Universidade Federal de Viçosa

V

Viviana Ruiz-Gutiérrez

Cornell Lab of Ornithology, Cornell University

L

Luís Fábio Silveira

Museu de Zoologia da Universidade de São Paulo

P

Philip C Stouffer

School of Renewable Natural Resources, Louisiana State University AgCenter and Louisiana State University

J

John W. Terborgh

Department of Biology, University of Florida

A

Alexandre Uezu

Instituto de Pesquisas Ecológicas

Y

Yanping Wang

Zhejiang Engineering Research Center for Biological Control of Crop Pathogens and Insects, Zhejiang Key Laboratory of Biology and Ecological Regulation of Crop Pathogens and Insects, Ministry of Agriculture and Rural Affairs Key Lab of Molecular Biology of Crop Pathogens and Insect Pests, State Key Lab of Rice Biology and Breeding, Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University

R

Robyn Wethered

Cossypha Ecological

D

Ding Li Yong

BirdLife International (Asia), Tanglin International Centre

C

Carlos A. Peres