Harnessing the CD2 axis to broaden and enhance the efficacy of CAR T-cell therapies

A Alberto Carturan (Lymphoma Program, Abramson Cancer Center, University of Pennsylvania, Philadelphia) M Mathew G. Angelos (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) P Puneeth Guruprasad R Ruchi P. Patel (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) R Raymone Pajarillo (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) A Andrew Lee D David Espie (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) Y Yunlin Zhang Y Yi-Hao Chiang (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) W Wei Xie J Jesse L. Rodriguez (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) J Jaryse Harris (6Division of Hematopathology, University of Pennsylvania, Philadelphia, PA) P Pooja Devi (Department of Pathology and Laboratory Medicine, Hospital of the University of Pennsylvania, Philadelphia) O Olabisi I. Afolayan-Oloye (6Division of Hematopathology, University of Pennsylvania, Philadelphia, PA) J Jason Xu J Jonathan H. Sussman O Omar Elghawy (2Division of Hematology and Oncology, University of Pennsylvania, Philadelphia, PA) A Austin Yang (7Division of Oncology, Children’s Hospital of Philadelphia, Philadelphia, PA) A Adam Barsouk (2Division of Hematology and Oncology, University of Pennsylvania, Philadelphia, PA) J Jong Hyun Cho C Carolyn E. Shaw (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) E Ekta Singh O Ositadimma Ugwuanyi (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) L Luca Paruzzo F Federico Stella (Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia) S Shan Liu S Siena Nason (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) A Antonio Imparato A Antonia Rotolo (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) J Jean Lemoine (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) D David M. Barrett A Avery Posey (1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA) A Alain H. Rook (10Department of Dermatology, University of Pennsylvania, Philadelphia, PA) V Vinodh Pillai A Adam Bagg (6Division of Hematopathology, University of Pennsylvania, Philadelphia, PA) S Stefano A. Pileri (12Division of Hematopathology, Istituto Europeo di Oncologia, Milan, Italy) D Dongfang Liu K Kai Tan S Stephen J. Schuster (Lymphoma Program, Abramson Cancer Center, Division of Hematology Oncology, Department of Medicine, University of Pennsylvania) D David T. Teachey P Patrizia Porazzi (Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia) M Marco Ruella

Abstract

Abstract Patients with T-cell lymphomas and leukemias have overall poor outcomes because of the lack of targeted and effective treatments, particularly in the relapsed and refractory settings. The development of chimeric antigen receptor (CAR) T cells against T-cell neoplasms is limited by a lack of discriminating T-cell antigens that allow for effective antitumor responses while preventing CAR T-cell fratricide. We hypothesized that targeting CD2, a pan–T-cell antigen, with anti-CD2 CAR T cells engineered to lack CD2 expression (CART2) would improve manufacturability and preclinical efficacy. Optimized CD2-knockout CART2 cells, generated using CRISPR-Cas9, eradicated primary patient-derived CD2+ hematological neoplasms in vitro and in vivo, secreted effector cytokines, and exhibited adequate proliferative capacity. Nevertheless, CD2 has a key costimulatory function, and its deletion could lead to CAR T-cell dysfunction. Therefore, we tested the role of the CD2:CD58 axis in CAR T cells, using the anti-CD19 CART models. We demonstrate that CD2 loss attenuates CART19 efficacy by reducing avidity for tumor antigen, costimulation, and ultimately in vivo activity. Analogously, we show that tumor CD58 loss reduces CART19 efficacy. To overcome this issue, we developed a novel programmed cell death protein 1 (PD-1):CD2 switch receptor that rescues intracellular CD2 signaling, particularly when programmed death-ligand 1 is engaged, thereby improving in vivo outcomes. Collectively, we studied the role of CD2 both as a target for CAR T-cell therapy and as a critical costimulatory protein, whose signaling can be rescued using the PD-1:CD2 switch receptor. This receptor can be incorporated into CAR T cells and provides an effective strategy to overcome CD2-signaling deficiencies.

Article Details

Journal Blood
Volume / Issue Vol. 147, Issue 16
Published April 16, 2026
Pages 1842-1856
ISSN 0006-4971
Publisher Elsevier BV

Journal Info

Blood

Elsevier BV

ISSN: 0006-4971 Health Sciences

Authors (42)

A

Alberto Carturan

Lymphoma Program, Abramson Cancer Center, University of Pennsylvania, Philadelphia

M

Mathew G. Angelos

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

P

Puneeth Guruprasad

R

Ruchi P. Patel

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

R

Raymone Pajarillo

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

A

Andrew Lee

D

David Espie

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

Y

Yunlin Zhang

Y

Yi-Hao Chiang

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

W

Wei Xie

J

Jesse L. Rodriguez

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

J

Jaryse Harris

6Division of Hematopathology, University of Pennsylvania, Philadelphia, PA

P

Pooja Devi

Department of Pathology and Laboratory Medicine, Hospital of the University of Pennsylvania, Philadelphia

O

Olabisi I. Afolayan-Oloye

6Division of Hematopathology, University of Pennsylvania, Philadelphia, PA

J

Jason Xu

J

Jonathan H. Sussman

O

Omar Elghawy

2Division of Hematology and Oncology, University of Pennsylvania, Philadelphia, PA

A

Austin Yang

7Division of Oncology, Children’s Hospital of Philadelphia, Philadelphia, PA

A

Adam Barsouk

2Division of Hematology and Oncology, University of Pennsylvania, Philadelphia, PA

J

Jong Hyun Cho

C

Carolyn E. Shaw

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

E

Ekta Singh

O

Ositadimma Ugwuanyi

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

L

Luca Paruzzo

F

Federico Stella

Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia

S

Shan Liu

S

Siena Nason

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

A

Antonio Imparato

A

Antonia Rotolo

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

J

Jean Lemoine

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

D

David M. Barrett

A

Avery Posey

1Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia, PA

A

Alain H. Rook

10Department of Dermatology, University of Pennsylvania, Philadelphia, PA

V

Vinodh Pillai

A

Adam Bagg

6Division of Hematopathology, University of Pennsylvania, Philadelphia, PA

S

Stefano A. Pileri

12Division of Hematopathology, Istituto Europeo di Oncologia, Milan, Italy

D

Dongfang Liu

K

Kai Tan

S

Stephen J. Schuster

Lymphoma Program, Abramson Cancer Center, Division of Hematology Oncology, Department of Medicine, University of Pennsylvania

D

David T. Teachey

P

Patrizia Porazzi

Center for Cellular Immunotherapies, University of Pennsylvania, Philadelphia

M

Marco Ruella