European beech reproduction is not reduced by drought, including the 2003, 2018, and 2022 extremes

J Jakub Szymkowiak (Forest Biology Center) M Michał Bogdziewicz (Forest Biology Center) D Dave Kelly (School of Biological Sciences) J Jessie Foest (Forest Biology Center) S Sabine Braun (Institute for Applied Plant Biology) B Burkhard Beudert (Department of Conservation and Research, Bavarian Forest National Park) F Francesco Chianucci (Council for Agricultural Research and Economics (CREA)—Research Centre for Forestry and Wood) A Andrea Cutini (Consiglio per la ricerca in agricoltura e l’analisi dell’economia agraria (CREA), Centro di Ricerca Foreste e Legno) R Rachel Gaulton (Fera Science Ltd., York Biotech Campus) G Georg Gratzer (Department of Ecosystem Management, Climate and Biodiversity, Institute of Forest Ecology, BOKU University) A Angelika Kölbl (Department of Conservation and Research, Bavarian Forest National Park) G Georges Kunstler (Université Grenoble Alpes, Institut national de recherche pour l’agriculture, l’alimentation et l’environnement (INRAE), Laboratoire EcoSystémes et Sociétés En Montagne (LESSEM)) J Jonathan G. A. Lageard (Department of Natural Sciences, Manchester Metropolitan University) H Henning Meesenburg (Northwest German Forest Research Institute, Department of Environmental Control, Section Intensive Environmental Monitoring) F Francesco Mezzavilla (Via Malviste 4) M Martina Mund (Forestry Research and Competence Centre Gotha) A Anita Nussbaumer (Swiss Federal Institute for Forest, Snow and Landscape Research WSL, Land Change Science) M Mario B. Pesendorfer (Department of Ecosystem Management, Climate and Biodiversity, Institute of Forest Ecology, BOKU University) W Wolfgang Schmidt A Anne Thimonier (Swiss Federal Institute for Forest, Snow and Landscape Research WSL, Land Change Science) P Peter A. Thomas (School of Life Sciences, Keele University) S Stanislav Vacek (Department of Silviculture, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Prague) Z Zdeněk Vacek (Department of Silviculture, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Prague) A Arne Verstraeten (Research Institute for Nature and Forest, Environment and Climate Unit) M Markus Wagner (Northwest German Forest Research Institute, Department of Environmental Control, Section Intensive Environmental Monitoring) A Andrew Hacket-Pain (Department of Geography and Planning)

Abstract

Climate change is intensifying drought stress in temperate forests, but its effects on tree reproduction, central to forest regeneration and migration capability, remain poorly understood. Here, we analyze 221 time series of beech ( Fagus sylvatica ) seed production across Europe to test whether drought reduces seed output. We isolate drought exposure during the flowering, pollination, and seed maturation phases of reproduction, and test for legacy effects on future reproduction. Seed production was not impaired by summer drought, and dry spring conditions were associated with increased output, likely via enhanced pollen dispersal. Thus, once initiated, beech reproduction is not reduced by drought, with no suppression of reproduction the following year. Reproduction was not reduced at the driest sites during exceptional European summer droughts in 2003, 2018, and 2022. Considered alongside prior evidence that drought suppresses forest growth and elevates mortality, these findings indicate that vital rates can respond in opposite directions to the same stressor. Such contrasts may sustain forest reproduction during heat–drought events yet shift demographic balance toward higher mortality and turnover as climatic extremes intensify.

Article Details

Volume / Issue Vol. 123, Issue 27
Published July 07, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (26)

J

Jakub Szymkowiak

Forest Biology Center

M

Michał Bogdziewicz

Forest Biology Center

D

Dave Kelly

School of Biological Sciences

J

Jessie Foest

Forest Biology Center

S

Sabine Braun

Institute for Applied Plant Biology

B

Burkhard Beudert

Department of Conservation and Research, Bavarian Forest National Park

F

Francesco Chianucci

Council for Agricultural Research and Economics (CREA)—Research Centre for Forestry and Wood

A

Andrea Cutini

Consiglio per la ricerca in agricoltura e l’analisi dell’economia agraria (CREA), Centro di Ricerca Foreste e Legno

R

Rachel Gaulton

Fera Science Ltd., York Biotech Campus

G

Georg Gratzer

Department of Ecosystem Management, Climate and Biodiversity, Institute of Forest Ecology, BOKU University

A

Angelika Kölbl

Department of Conservation and Research, Bavarian Forest National Park

G

Georges Kunstler

Université Grenoble Alpes, Institut national de recherche pour l’agriculture, l’alimentation et l’environnement (INRAE), Laboratoire EcoSystémes et Sociétés En Montagne (LESSEM)

J

Jonathan G. A. Lageard

Department of Natural Sciences, Manchester Metropolitan University

H

Henning Meesenburg

Northwest German Forest Research Institute, Department of Environmental Control, Section Intensive Environmental Monitoring

F

Francesco Mezzavilla

Via Malviste 4

M

Martina Mund

Forestry Research and Competence Centre Gotha

A

Anita Nussbaumer

Swiss Federal Institute for Forest, Snow and Landscape Research WSL, Land Change Science

M

Mario B. Pesendorfer

Department of Ecosystem Management, Climate and Biodiversity, Institute of Forest Ecology, BOKU University

W

Wolfgang Schmidt

A

Anne Thimonier

Swiss Federal Institute for Forest, Snow and Landscape Research WSL, Land Change Science

P

Peter A. Thomas

School of Life Sciences, Keele University

S

Stanislav Vacek

Department of Silviculture, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Prague

Z

Zdeněk Vacek

Department of Silviculture, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Prague

A

Arne Verstraeten

Research Institute for Nature and Forest, Environment and Climate Unit

M

Markus Wagner

Northwest German Forest Research Institute, Department of Environmental Control, Section Intensive Environmental Monitoring

A

Andrew Hacket-Pain

Department of Geography and Planning