Epidemiology, clinical features, and molecular basis of <i>TTMV</i> :: <i>RARA</i> -driven acute promyelocytic leukemia

X Xiaosu Zhou (2Beijing Lu Daopei Institute of Hematology, Beijing, China) J Jiaqi Chen Y Yan-Lai Tang (5Department of Pediatrics, The First Affiliated Hospital, Sun Yat-sen University, Guangzhou, China) H Haibo Sun Q Qihui Chen (State Key Laboratory of Structural Chemistry, Fujian Institute of Research on the Structure of Matter) X Xue Chen Y Yang Zhang J Jiancheng Fang (Institute of Large-Scale Scientific Facility, School of Instrumentation Science and Opto-Electronics Engineering, Beihang University) P Panxiang Cao (1Hebei Yanda Lu Daopei Hospital, Langfang, China) X Xiaoli Ma (Institute of Physics, Chinese Academy of Sciences) L Li Chen Y YangYang Xie (Chinese Academy of Sciences Key Laboratory of Nutrition, Metabolism and Food Safety, Shanghai Institute of Nutrition and Health, University of Chinese Academy of Sciences, Chinese Academy of Sciences) L Lijun Wen L Li-Bin Huang Z Zhong Fan (1Univeristy of Texas Health Science Center at Houston, Houston, United States) J Jiacheng Lou (7Shanghai Institute of Hematology, State Key Laboratory of Medical Genomics, National Research Center for Translational Medicine at Shanghai, Ruijin Hospital Affiliated with Shanghai Jiao Tong University School of Medicine, Shanghai, China) W Wenning Xu (11Department of Hematology and Lymphoma, The First People's Hospital of Foshan, Foshan, China) N Nan Zhang J Jing Li Y Yu Luo (Macau Centre for Research and Development in Chinese Medicine, State Key Laboratory of Mechanism and Quality of Chinese Medicine, Institute of Chinese Medical Sciences) Y Yan Dai H Huan Du (State Key Laboratory of Chemo and Biosensing, College of Chemistry and Chemical Engineering) K Kun Yang Z Zhixu He (19Department of Pediatrics, Affiliated Hospital of Guizhou Medical University, Guiyang, China) X Xiaoyan Yang L Linya Wang (1Peking University People's Hospital, Beijing, China) B Bing Liu D Danna Lin (23Department of Pediatric Hematology, Zhujiang Hospital, Southern Medical University, Guangzhou, China) Y Yajie Zhang (Center for Carbon-Based Electronics and Key Laboratory for the Physics and Chemistry of Nanodevices, School of Electronics) F Fang Xu (Key Laboratory of Optoelectronic Chemical Materials and Devices (Ministry of Education), School of Optoelectronic Materials and Technology) M Min Zhang Y Yang Wang S Shu Yan H Hua Nan J Jing Zhang L Li Xu X Xiujuan Ma R Ruijie Tang F Fang Wang L Lili Yuan (1Hebei Yanda Lu Daopei Hospital, Langfang, Langfang, China) M Ming Liu L Lei Wang Q Qisheng Wu L Lina Zhang S Siyuan Liu (Sydney Dental School, Faculty of Medicine and Health, Charles Perkins Centre) C Chengcheng Yan (2Tianjin Kingmed Diagnostics Laboratory Co.Ltd, Tianjin, China., Tianjin, China) J Jianfeng Zhou F Fanyong Lyu (28Department of Hematology, Beijing Lu Daopei Hospital, Beijing, China) P Ping Wu (Department of Neurobiology, University of Texas Medical Branch) T Tong Wang X Xian Zhang (State Key Laboratory of Analytical Chemistry for Life Science, School of Chemistry) Y Yue Lu D Deyan Liu (1Hebei Yanda Lu Daopei Hospital, Langfang, China) J Junfang Yang (1Hebei Yanda Lu Daopei Hospital, Langfang, China) M Min Xiong Y Yanli Zhao (School of Chemistry, Chemical Engineering and Biotechnology) G Gaowei Fan X Xia Xiao F Futian Ma (16Department of Hematology, Hebei Children’s Hospital, Shijiazhuang, China) L Lihua Yang X Xue-Qun Luo (5Department of Pediatrics, The First Affiliated Hospital, Sun Yat-sen University, Guangzhou, China) J Junmin Li S Shuhong Shen (1Department of Hematology/Oncology, Shanghai Children’s Medical Center, Shanghai Jiao Tong University School of Medicine, National Health Committee Key Laboratory of Pediatric Hematology and Oncology, Shanghai, China) H Huyong Zheng (1Beijing Children's Hospital, Capital Medical University, Beijing, China) S Suning Chen H Hongwei Chen (State Key Laboratory of Coordination Chemistry, Chemistry and Biomedicine Innovation Center (ChemBIC), ChemBioMed Interdisciplinary Research Center at Nanjing University, School of Chemistry) J Jianhong Zhao (Jining No.1 People’s Hospital, Jining, China) Z Zhanglin Zhang (4Department of Blood Transfusion, The First Affiliated Hospital of Nanchang University, Jiangxi Key Laboratory of transfusion, Institute of Transfusion, Jiangxi Academy of Clinical Medical Sciences, Jiangxi Medical College, Nanchang University, Nanchang, China) K Kankan Wang P Peihua Lu (1Hebei Yanda Lu Daopei Hospital, Langfang, China) H Hongxing Liu (1Hebei Yanda Lu Daopei Hospital, Langfang, China)

Abstract

Abstract Integration of torque teno mini virus (TTMV) generating the TTMV::RARA (retinoic acid receptor α) fusion represents a newly recognized subtype of acute promyelocytic leukemia (APL) that merits detailed investigation. We present, to our knowledge, the first comprehensive characterization of its epidemiologic profile, clinical presentation, virologic characteristics, and underlying molecular mechanisms. Our findings indicate that TTMV::RARA is more prevalent in pediatric patients and represents the second most common retinoic acid receptor fusion after PML::RARA. Affected patients exhibit a high incidence of extramedullary involvement, particularly myeloid sarcoma. Cytogenetic abnormalities involving i(17)(q10) or 7q22 were identified in 52.0% of cases, largely in a mutually exclusive manner. Co-occurring mutations in epigenetic regulators were present in 76.9% of patients. Although most patients achieved initial remission, relapse was common and associated with rapid acquisition of all-trans retinoic acid (ATRA)–resistant mutation and secondary chemoresistance. Venetoclax-containing regimens demonstrated encouraging clinical efficacy. Phylogenetic analysis indicated that patient-derived TTMV strains clustered into a distinct clade. TTMV integration consistently occurred within RARA intron 2, involving a consensus fragment of 510 to 610 base pairs encompassing the viral promoter and open reading frame 2 (ORF2) N terminus, likely mediated by microhomology-driven recombination. Tandem RUNX1-binding motifs within the integrated viral promoter may underlie the myelotropism of these TTMV strains and facilitate transcriptional activation of TTMV::RARA. The chimeric protein retains at least the first 56 N-terminal residues of ORF2 and remains transcriptionally responsive to pharmacological concentrations of ATRA. These findings establish TTMV::RARA-APL as a distinct leukemia entity, laying the foundation for future studies on virus-mediated leukemogenesis and therapeutic strategies.

Article Details

Journal Blood
Volume / Issue Vol. 146, Issue 18
Published October 30, 2025
Pages 2229-2243
ISSN 0006-4971
Publisher Elsevier BV

Journal Info

Blood

Elsevier BV

ISSN: 0006-4971 Health Sciences

Authors (71)

X

Xiaosu Zhou

2Beijing Lu Daopei Institute of Hematology, Beijing, China

J

Jiaqi Chen

Y

Yan-Lai Tang

5Department of Pediatrics, The First Affiliated Hospital, Sun Yat-sen University, Guangzhou, China

H

Haibo Sun

Q

Qihui Chen

State Key Laboratory of Structural Chemistry, Fujian Institute of Research on the Structure of Matter

X

Xue Chen

Y

Yang Zhang

J

Jiancheng Fang

Institute of Large-Scale Scientific Facility, School of Instrumentation Science and Opto-Electronics Engineering, Beihang University

P

Panxiang Cao

1Hebei Yanda Lu Daopei Hospital, Langfang, China

X

Xiaoli Ma

Institute of Physics, Chinese Academy of Sciences

L

Li Chen

Y

YangYang Xie

Chinese Academy of Sciences Key Laboratory of Nutrition, Metabolism and Food Safety, Shanghai Institute of Nutrition and Health, University of Chinese Academy of Sciences, Chinese Academy of Sciences

L

Lijun Wen

L

Li-Bin Huang

Z

Zhong Fan

1Univeristy of Texas Health Science Center at Houston, Houston, United States

J

Jiacheng Lou

7Shanghai Institute of Hematology, State Key Laboratory of Medical Genomics, National Research Center for Translational Medicine at Shanghai, Ruijin Hospital Affiliated with Shanghai Jiao Tong University School of Medicine, Shanghai, China

W

Wenning Xu

11Department of Hematology and Lymphoma, The First People's Hospital of Foshan, Foshan, China

N

Nan Zhang

J

Jing Li

Y

Yu Luo

Macau Centre for Research and Development in Chinese Medicine, State Key Laboratory of Mechanism and Quality of Chinese Medicine, Institute of Chinese Medical Sciences

Y

Yan Dai

H

Huan Du

State Key Laboratory of Chemo and Biosensing, College of Chemistry and Chemical Engineering

K

Kun Yang

Z

Zhixu He

19Department of Pediatrics, Affiliated Hospital of Guizhou Medical University, Guiyang, China

X

Xiaoyan Yang

L

Linya Wang

1Peking University People's Hospital, Beijing, China

B

Bing Liu

D

Danna Lin

23Department of Pediatric Hematology, Zhujiang Hospital, Southern Medical University, Guangzhou, China

Y

Yajie Zhang

Center for Carbon-Based Electronics and Key Laboratory for the Physics and Chemistry of Nanodevices, School of Electronics

F

Fang Xu

Key Laboratory of Optoelectronic Chemical Materials and Devices (Ministry of Education), School of Optoelectronic Materials and Technology

M

Min Zhang

Y

Yang Wang

S

Shu Yan

H

Hua Nan

J

Jing Zhang

L

Li Xu

X

Xiujuan Ma

R

Ruijie Tang

F

Fang Wang

L

Lili Yuan

1Hebei Yanda Lu Daopei Hospital, Langfang, Langfang, China

M

Ming Liu

L

Lei Wang

Q

Qisheng Wu

L

Lina Zhang

S

Siyuan Liu

Sydney Dental School, Faculty of Medicine and Health, Charles Perkins Centre

C

Chengcheng Yan

2Tianjin Kingmed Diagnostics Laboratory Co.Ltd, Tianjin, China., Tianjin, China

J

Jianfeng Zhou

F

Fanyong Lyu

28Department of Hematology, Beijing Lu Daopei Hospital, Beijing, China

P

Ping Wu

Department of Neurobiology, University of Texas Medical Branch

T

Tong Wang

X

Xian Zhang

State Key Laboratory of Analytical Chemistry for Life Science, School of Chemistry

Y

Yue Lu

D

Deyan Liu

1Hebei Yanda Lu Daopei Hospital, Langfang, China

J

Junfang Yang

1Hebei Yanda Lu Daopei Hospital, Langfang, China

M

Min Xiong

Y

Yanli Zhao

School of Chemistry, Chemical Engineering and Biotechnology

G

Gaowei Fan

X

Xia Xiao

F

Futian Ma

16Department of Hematology, Hebei Children’s Hospital, Shijiazhuang, China

L

Lihua Yang

X

Xue-Qun Luo

5Department of Pediatrics, The First Affiliated Hospital, Sun Yat-sen University, Guangzhou, China

J

Junmin Li

S

Shuhong Shen

1Department of Hematology/Oncology, Shanghai Children’s Medical Center, Shanghai Jiao Tong University School of Medicine, National Health Committee Key Laboratory of Pediatric Hematology and Oncology, Shanghai, China

H

Huyong Zheng

1Beijing Children's Hospital, Capital Medical University, Beijing, China

S

Suning Chen

H

Hongwei Chen

State Key Laboratory of Coordination Chemistry, Chemistry and Biomedicine Innovation Center (ChemBIC), ChemBioMed Interdisciplinary Research Center at Nanjing University, School of Chemistry

J

Jianhong Zhao

Jining No.1 People’s Hospital, Jining, China

Z

Zhanglin Zhang

4Department of Blood Transfusion, The First Affiliated Hospital of Nanchang University, Jiangxi Key Laboratory of transfusion, Institute of Transfusion, Jiangxi Academy of Clinical Medical Sciences, Jiangxi Medical College, Nanchang University, Nanchang, China

K

Kankan Wang

P

Peihua Lu

1Hebei Yanda Lu Daopei Hospital, Langfang, China

H

Hongxing Liu

1Hebei Yanda Lu Daopei Hospital, Langfang, China