Decoding antibody response to MERS-CoV in wild dromedary camels

F Faten Idoudi (Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar) R Ruoke Wang (Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University) L Long Tian (Key Laboratory of Rare Earth Resource Utilization, Changchun Institute of Applied Chemistry) Z Ziqing Yang (Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University) K Kenn Ka-Heng Chik (State Key Laboratory of Emerging Infectious Diseases, Carol Yu Centre for Infection, Department of Microbiology, Li Ka Shing Faculty of Medicine, The University of Hong Kong) P Peng Chen R Rahma Benabderrazek (Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar) L Lili Fu (Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University) R Ruijie Tan (Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University) S Sayda Dhaouadi (Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar) Z Zakaria Benlasfar (Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar) M Mahmoud Somia (Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar) Q Qi Zhang X Xuanling Shi (Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University) J Jasper Fuk-Woo Chan (Department of Microbiology, School of Clinical Medicine, Li Ka Shing Faculty of Medicine, The University of Hong Kong) K Kwok-Yung Yuen (Department of Microbiology, Li Ka Shing Faculty of Medicine, The University of Hong Kong) X Xinquan Wang (The Ministry of Education Key Laboratory of Protein Science) B Balkiss Bouhaouala-Zahar (Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar) L Linqi Zhang (Pandemic Research Alliance Unit at the University of Hong Kong)

Abstract

Wild dromedary camels in the Arabian Peninsula and Africa have harbored antibodies against Middle East Respiratory Syndrome Coronavirus (MERS-CoV) for decades, predating zoonotic spillover to humans. However, the potency, specificity, and structural characteristics of these antibodies remain poorly understood. Here, we characterize the antibody responses of naturally infected wild dromedary camels in Tunisia, a MERS-CoV-endemic region. Plasma antibodies from nine camels exhibited variable neutralizing activity, generally increasing with age, and were largely autologous, with minimal cross-reactivity to SARS-CoV-1 or SARS-CoV-2. From a VHH antibody library derived from the peripheral blood mononuclear cells (PBMCs) of a single camel (D17), we identified 34 unique sequences with previously unreported germline origins and unusually long complementarity-determining region 3 (CDR3) sequences. Eight representative VHHs, expressed as human Fc fusions, displayed high-affinity binding to the MERS-CoV receptor-binding domain (RBD) and broad neutralization to RBD mutants (IC 50 : 1.05 to 9.55 ng/mL). Crystal structural analysis revealed distinct neutralization mechanisms: VHH-227 fully blocked DPP4 binding, achieving complete neutralization, while VHH-T71, with partial neutralization (~80%), targeted the RBD core subdomain. This study provides comprehensive characterization of wild dromedary antibody responses, identifying novel nanobodies (Nbs) with broad and potent neutralization to naturally occurring RBD mutants. These findings offer insights into camel immunity and highlight promising candidates for MERS-CoV prophylactic and therapeutic development.

Article Details

Volume / Issue Vol. 123, Issue 7
Published February 17, 2026
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (19)

F

Faten Idoudi

Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar

R

Ruoke Wang

Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University

L

Long Tian

Key Laboratory of Rare Earth Resource Utilization, Changchun Institute of Applied Chemistry

Z

Ziqing Yang

Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University

K

Kenn Ka-Heng Chik

State Key Laboratory of Emerging Infectious Diseases, Carol Yu Centre for Infection, Department of Microbiology, Li Ka Shing Faculty of Medicine, The University of Hong Kong

P

Peng Chen

R

Rahma Benabderrazek

Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar

L

Lili Fu

Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University

R

Ruijie Tan

Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University

S

Sayda Dhaouadi

Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar

Z

Zakaria Benlasfar

Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar

M

Mahmoud Somia

Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar

Q

Qi Zhang

X

Xuanling Shi

Comprehensive AIDS Research Center, Pandemic Research Alliance Unit, Center for Infection Biology, School of Basic Medical Sciences, Tsinghua University

J

Jasper Fuk-Woo Chan

Department of Microbiology, School of Clinical Medicine, Li Ka Shing Faculty of Medicine, The University of Hong Kong

K

Kwok-Yung Yuen

Department of Microbiology, Li Ka Shing Faculty of Medicine, The University of Hong Kong

X

Xinquan Wang

The Ministry of Education Key Laboratory of Protein Science

B

Balkiss Bouhaouala-Zahar

Pasteur Institute of Tunis, Laboratory of Biomolecules, Venoms and Theranostic Applications (LR20IPT01), NanoBioMedika research Team, University Tunis El Manar

L

Linqi Zhang

Pandemic Research Alliance Unit at the University of Hong Kong