ARRDC4-mediated glycolysis enhances innate immunity to influenza A virus through fructose-1,6-bisphosphate

Y Yuhan Li Z Zhen Wang J Jie Wang (State Key Laboratory of Molecular Oncology, Beijing Key Laboratory, CAMS Key Laboratory of Translational Research on Lung Cancer, Department of Medical Oncology Cancer Hospital, Chinese Academy of Medical Sciences Beijing China) Z Zhimin Jiang (Guangxi Key Laboratory of Drug Discovery and Optimization, School of Pharmacy) M Mingyue Chen (State Key Laboratory of Veterinary Public Health and Safety, Department of Preventive Veterinary Medicine, College of Veterinary Medicine, China Agricultural University) H Hui Ai (State Key Laboratory of Veterinary Public Health and Safety, Department of Preventive Veterinary Medicine, College of Veterinary Medicine, China Agricultural University) C Chao Ma Q Qi Tong (National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University) L Litao Liu (National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University) T Tony Velkov (Department of Pharmacology, Biodiscovery Institute, Monash University) H Honglei Sun (National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University) J Juan Pu (National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University) J Jinhua Liu (National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University) C Chongshan Dai (State Key Laboratory of Veterinary Public Health and Safety, Department of Preventive Veterinary Medicine, College of Veterinary Medicine, China Agricultural University) Y Yipeng Sun

Abstract

Glucose metabolism impacts the innate immune response against viral infection. However, the key enzymes or the natural products and mechanisms involved are not well elucidated. Here, we found that arrestin domain containing 4 (ARRDC4), a critical regulator of glucose metabolism, senses influenza A virus (IAV) infection by interacting with viral PA protein. Upregulated ARRDC4 increases the enzymatic activity of phosphofructokinase, muscle type (PFKM) via binding its His298 site to promote the production of the metabolite fructose-1,6-bisphosphate (FBP). Consequently, FBP inhibits the K48-linked ubiquitination degradation of HSP90β, subsequently enhances its interaction with IKKβ and IKKε, and enhances NF-κB- and IRF7-mediated antiviral innate immunity, respectively. Importantly, FBP supplementation enhanced IFN-β-mediated antiviral innate immunity in vitro and in vivo. Our findings highlight a unique immunometabolic regulatory mechanism in which ARRDC4 senses IAV infection and regulates antiviral innate immunity through the PFKM-FBP metabolic axis and provide a strategy for manipulating FBP-related metabolism to treat viral infection.

Article Details

Volume / Issue Vol. 122, Issue 35
Published September 02, 2025
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (15)

Y

Yuhan Li

Z

Zhen Wang

J

Jie Wang

State Key Laboratory of Molecular Oncology, Beijing Key Laboratory, CAMS Key Laboratory of Translational Research on Lung Cancer, Department of Medical Oncology Cancer Hospital, Chinese Academy of Medical Sciences Beijing China

Z

Zhimin Jiang

Guangxi Key Laboratory of Drug Discovery and Optimization, School of Pharmacy

M

Mingyue Chen

State Key Laboratory of Veterinary Public Health and Safety, Department of Preventive Veterinary Medicine, College of Veterinary Medicine, China Agricultural University

H

Hui Ai

State Key Laboratory of Veterinary Public Health and Safety, Department of Preventive Veterinary Medicine, College of Veterinary Medicine, China Agricultural University

C

Chao Ma

Q

Qi Tong

National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University

L

Litao Liu

National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University

T

Tony Velkov

Department of Pharmacology, Biodiscovery Institute, Monash University

H

Honglei Sun

National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University

J

Juan Pu

National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University

J

Jinhua Liu

National Key Laboratory of Veterinary Public Health and Safety, Key Laboratory for Prevention and Control of Avian Influenza and Other Major Poultry Diseases of the Ministry of Agriculture and Rural Affairs, College of Veterinary Medicine, China Agricultural University

C

Chongshan Dai

State Key Laboratory of Veterinary Public Health and Safety, Department of Preventive Veterinary Medicine, College of Veterinary Medicine, China Agricultural University

Y

Yipeng Sun