Alternative cGAS signaling promotes herpes simplex encephalitis

L Liraz Shmuel-Galia (Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School) Z Zhaozhao Jiang (Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School) L Laurel Stine (Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School) S Sara Cahill (Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School) S Sze-Ling Ng (Innate Immunity Research Unit, GlaxoSmithKline) R Ruth Wilson (Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School) R Richard Kumaran Kandasamy (Centre of Molecular Inflammation Research, and Department of Clinical and Molecular Medicine, Norwegian University of Science and Technology) E Evelyn A. Kurt-Jones (Division of Infectious Diseases and Immunology, Department of Medicine, University of Massachusetts Chan Medical) J Joshi M. Ramanjulu (Innate Immunity Research Unit, GlaxoSmithKline) J John Bertin (Immunology and Inflammation Therapeutic Area, Sanofi) V Viera Kasparcova (Innate Immunity Research Unit, GlaxoSmithKline) G G. Scott Pesiridis (Innate Immunity Research Unit, GlaxoSmithKline) K Katherine A. Fitzgerald (Department of Medicine, University of Massachusetts Chan Medical School) F Fiachra Humphries

Abstract

During infection, foreign DNA is sensed by cyclic GMP-AMP synthase (cGAS) leading to the production of cGAMP, STING-dependent type I interferon and proinflammatory cytokine expression, and autophagy. To prevent a response to self-DNA, cGAS activity is tightly regulated. Dysregulation of cGAS underpins interferonopathies, such as Aicardi–Goutières syndrome, as well as Lupus and neurodegenerative diseases like Parkinson’s disease. Thus, cGAS and its product cGAMP are therapeutic targets. However, if cGAS functions independently of cGAMP signaling is undefined. Here, we identified an alternative signaling pathway that cGAS engages independent of cGAMP synthesis. We demonstrate that alternative cGAS signaling promotes hyperexpression of CXCL1 and enhanced neutrophil recruitment that facilitates viral dissemination during herpes simplex encephalitis. Our study reports of an alternative cGAS response independent of cGAMP, highlighting a previously uncharacterized scaffold function for cGAS.

Article Details

Volume / Issue Vol. 122, Issue 22
Published June 03, 2025
ISSN 0027-8424
Publisher National Academy of Sciences

Authors (14)

L

Liraz Shmuel-Galia

Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School

Z

Zhaozhao Jiang

Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School

L

Laurel Stine

Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School

S

Sara Cahill

Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School

S

Sze-Ling Ng

Innate Immunity Research Unit, GlaxoSmithKline

R

Ruth Wilson

Division of Innate Immunity, Department of Medicine, University of Massachusetts Chan Medical School

R

Richard Kumaran Kandasamy

Centre of Molecular Inflammation Research, and Department of Clinical and Molecular Medicine, Norwegian University of Science and Technology

E

Evelyn A. Kurt-Jones

Division of Infectious Diseases and Immunology, Department of Medicine, University of Massachusetts Chan Medical

J

Joshi M. Ramanjulu

Innate Immunity Research Unit, GlaxoSmithKline

J

John Bertin

Immunology and Inflammation Therapeutic Area, Sanofi

V

Viera Kasparcova

Innate Immunity Research Unit, GlaxoSmithKline

G

G. Scott Pesiridis

Innate Immunity Research Unit, GlaxoSmithKline

K

Katherine A. Fitzgerald

Department of Medicine, University of Massachusetts Chan Medical School

F

Fiachra Humphries