Xinyang Wang, Yixiong Hu, Yajiao Wen, Miaomiao Shen, Pu Chen, Song Xiang. "SHPRH together with the Ube2D family of enzymes directly ubiquitinates PCNA at Lys164 in vitro." PLoS ONE (2026). https://doi.org/10.1371/journal.pone.0347227